The paper has benefited from discussions with Philippe Arbogast, Olivia Martius, Guillaume Lapeyre, and Alain Joly and from constructive comments of three anonymous reviewers. The authors thank Marie Boisserie for her English corrections in an early version of the manuscript. This work has been partially funded by a CNRS/INSU/LEFE/IDAO grant and an ANR grant (ANR-06-JCJC-133-01).
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