This research was sponsored by National Science Foundation Grants ATM-9616728 and ATM-9627890. This is Journal Paper No. J-18448 of the Iowa Iowa Agriculture and Home Economics Experiment Station, Ames, Iowa, Project No. 3245, and supported by Hatch Act and State of Iowa. We thank Dennis Todey for his assistance with figure preparation. This manuscript was improved significantly by the comments of two anonymous reviewers.
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