Funding for this work has been provided by the National Park Service (NPS) and the National Biological Survey (NBS) Grants CEGR-R92-0193 and COLR-R92-0204, Environmental Protection Agency (EPA) Grant R824993-01-0, and NASA Grants NAG8-1511 and NAG5-4646. The authors would like to acknowledge the anonymous reviewers for their excellent suggestions. This paper forms part of the first author’s Ph.D. dissertation.
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