The study was supported by NSF Grants ATM-9909650 and ATM- 9911417, and the ISU Agronomy Endowment Fund. This is Journal Paper J-19316 of the Iowa Agricultural and Home Economics Experiment Station, Ames, Iowa, Project 3803, and supported by the Hatch Act and State of Iowa. Constructive comments by two anonymous reviewers improved the original version of this manuscript. Reatha Diedrichs prepared the manuscript.
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