1. Introduction
The stability properties of baroclinic zonal flows on the β plane have been long studied as a consequence of their relevance to the explanation of the development of wavelike disturbances in atmospheric flow whose wavefree state would be essentially zonal. Necessary conditions for instability, bounds on growth rates, and phase speeds of unstable normal modes are well known [see Pedlosky (1987) for a review]. Additional theorems speaking to the range of wavenumbers, in particular the maximum allowable wavenumber for instability for the modes, can also be developed in some cases (Pedlosky 1964). Particular examples of the normal modes of instability of such flows have formed the conceptual paradigm most researchers rely upon in thinking about the general stability properties of atmospheric and oceanic flows.
While such models are certainly pertinent to those oceanic flows that are largely zonal, the ocean provides examples of broad, baroclinic flows that deviate substantially from zonality. Indeed, in the eastern portions of the major subtropical gyres the flow, constrained by the presence of the eastern boundary of its basin, is more meridional than zonal. The shears of such Sverdrup interior flows are relatively weak compared to the western boundary currents but they contain substantial stores of available potential energy and it is natural to anticipate local instabilities growing on that potential energy stored in the east–west slope of the supporting density surfaces in the meridional shear.
If the flow were horizontally uniform and laterally unbounded, such currents would always be unstable. This is possible since disturbances consisting of plane waves with crests oriented zonally would release the potential energy of the current without suffering the stabilizing effects of β because the perturbation velocities would be strictly zonal. Such an idealized limit is, of course, never realized. The finite size of the gyre, the variation of the basic current, and the inevitable nonuniformity of initial conditions will always impose a zonal variation of the structure of the disturbance. It is natural then to ask what the role of the planetary vorticity gradient will be in the development of the instabilities of meridional currents of laterally finite extent.
Spall (1994, 2000) has addressed some of these issues in papers dealing with the variability of the circulation of the eastern North Atlantic. The emphasis in his papers cited above is the nonlinear development of disturbances from initial data, their eventual amplitude equilibration, and their effect on tracer fields such as salinity. The disturbances in his numerical calculation are permitted to develop and leave the domain of the flow through the device of an absorbing layer near the western boundary of the region of calculation. The flow is strongly unstable and secondary instabilities spontaneously develop before the disturbances are able to evolve to anything resembling a normal mode structure.
Kamenkovich and Pedlosky (1996) examined the linear instability of nonzonal jets and showed that the combination of nonzonality and wave radiation could destabilize otherwise stable flows. Indeed, it is the nonzonality that is responsible for the radiation. The study concentrated on jets in unbounded regions but did not confront directly the case of purely meridional flow.
We believe it would be of value to understand the underlying normal mode structure of such flows, their characteristic growth rates, and propagation speeds as well as the parameter domain in which they would be observed and the relation of such modes to both the initial value problem and the subsequent nonlinear evolution and equilibration of the disturbances. The normal mode problem is an important element in understanding the basic physics of the instability of such flows, even when the individual modes only eventuate for very long times. The spectrum of the modes presents us with bounds on growth rates and useful information of preferred structures.
The present study forms a first step in that effort and presents an analysis of the normal modes of an idealized channel flow oriented in the north–south direction. This idealization is meant to substitute for the longitudinal variation a current of finite width would impose on the perturbation without the added complexity of dealing with both horizontal and vertical shear. The flow in this study has only vertical shear and the channel width can be of arbitrary size with respect to the deformation radius. This is, therefore, the Phillips model (1954) turned 90 degrees.
We find that in contrast to the zonal flow case there is no minimum shear required for instability. Similarly, we find that there is instability even for channel widths less than the deformation radius. The modal structure in the cross-channel direction is a strong function of the parameters and the mode's growth rate. We infer from the sum of our results a strong connection between the instability occurring at weak shears and the existence of Rossby normal modes although we have not been able to prove the connection conclusively. Although the growth rates of the cases with weak shears are small, there is no reason to believe that the resulting amplitudes in an equilibrated nonlinear state need be small. Indeed, the analysis of Shepherd (1988) shows how an a priori bound on the amplitude of the equilibrated state can be given in terms of the degree of supercriticality of the flow. Absent a threshold for instability, the supercriticality is always O(1) and it is natural to assume that the resulting equilibrated amplitudes will be consequently larger than in the zonal flow case. That will depend on the efficacy of the equilibration process, which itself may be weak for broad meridional flows, as Spall (2000) suggests; therefore, the existence of such weak-shear modes may be oceanographically very significant.
Section two describes the basic model and its formulation. The symmetry properties of the model when the layer thicknesses are equal are described, as well as the symmetry between northward and southward flowing currents. Section 3 gives a preliminary overview of our detailed calculation and provides a starting point for an analytical prediction of both the long-wave and short-wave cutoffs of the model. We also present in section 4 an asymptotic analysis of the instabilities for a very broad channel that illustrates the tendency for western intensification of the modes of instability. Section 5 is a detailed look at the modal calculations in which the phase speeds and growth rates for the modes are presented and contrasted with the zonal flow case. Section 6 briefly describes the model with unequal layer thicknesses in which certain symmetries of the basic model are broken. Section 7 describes the results of initial value calculations that provide a link between the modal theory and numerical calculations of Spall, at least in the early linear stages. We conclude in section 8 with a discussion of our results and their significance.
2. The model
We employ throughout the two-layer quasigeostrophic model on the β plane (Pedlosky 1987). Since the basic flow is nonzonal, this implies a potential vorticity (PV) source in each layer for which the basic flow is nonzonal. We will restrict attention to cases in which the basic flow is a uniform flow in only the upper of the two layers so that the PV forcing is only in the upper layer. The reader may think of it as a uniform wind stress curl acting on the upper layer alone.




It is important to note that the major physical difference between the problem of the instability of a meridional shear and the zonal equivalent is that, as can be seen from (2.3), the potential vorticity gradients associated with β and the potential vorticity gradient of the basic flow are not colinear. It is therefore impossible for the beta effect to completely determine the sign of the potential vorticity gradient of the ambient flow as seen by the perturbation field. This is the fundamental physical novelty of the meridional instability problem.

Try as we might, we have not been able, using the usual integral methods, to deduce a useful necessary condition for instability, that is, one that would give us a maximum value of β beyond which the flow would be definitely stable. Nor have we been able to deduce useful bounds on phase speeds and growth rates. The presence of the complex term in (2.7) has thwarted such attempts, and we have come to believe this is a reflection of the absence of a well-defined stability threshold, a result which is further suggested by our calculations that follow.

We will also find solutions in which the real part of s is identically zero so that the perturbation is moving exactly with the mean speed of the current. The two “modes” obtained in that case from the symmetry property are really the same mode but the invariance in that case implies that the phase lines in the two layers will be sloping in opposite directions.
Similarly, under a change in sign of the basic flow it is possible, using the invariant properties of the equations, to show that a solution with positive V implies the same solution for the same negative V with a change in sign of the real phase speed and an interchange of wave functions between the two layers with the same growth rate.
These precise symmetries are broken when the layer thicknesses are unequal, as will be discussed in section 6.
3. Overview of results and long- and short-wave cutoffs
a. Method of solution

This leads to a fourth-order polynomial equation for kj. For a given c, the roots can be found numerically or, since the quartic is analytically tractable, the solutions can be found directly from the standard solution for quartics (Abramowitz and Stegun 1972). For each root the ratio a1j/a2j is determined by either of (2.3a,b). Application of the four boundary conditions at the channel boundaries yields a set of four homogeneous algebraic equations whose determinant must vanish for nontrivial solutions. Only for the complex eigenvalues of the problem will the determinant vanish and this condition, implemented numerically, provides us with the eigenvalues and the eigenfunctions from (3.1). These solutions were checked by simply discretizing the ordinary differential equations (2.3a,b), which turns the problem into a large matrix problem
b. A preliminary sample of results
Before proceeding with further analytical results it is useful to examine in a preliminary way the results of the numerical calculations outlined above since it is those results that suggested the analytical approaches to be presented and justify what might seem to be arbitrary assumptions about the solutions.
Figure 1 shows the eigenvalues c for the case where L = 10, β = 0.5 and for equal layer thicknesses. The eigenvalues of the first two modes are shown. Starting at the right-hand side of the figure, at short waves, we see two branches of the first mode with real phase speeds symmetrically centered around the mean flow speed of c = 0.5. As the y wavenumber is reduced, the two branches coalesce at the phase speed c = 0.5. This is found to be a general birthplace for unstable modes. This occurs first at the point

c. Eigenfunctions
For the value of β in Figs. 1 and 2 we present in Fig. 3 the eigenfunctions in the channel at selected wavenumbers. The panel in the upper left-hand corner identifies the eigenvalue associated with each eigenfunction. Each panel in the rest of the figure is identified by a letter identifying the associated eigenvalue. Each such panel is doubled, the left-hand side representing the upper layer and the right-hand panel representing the lower layer.
It is important to bear in mind that the y wavenumber is O(1) and the channel width is L = 10, so for the purposes of spatial compression the aspect ratio has been artificially foreshortened. We see that unlike the zonal flow case the horizontal structure of the eigenfunctions strongly varies in parameter space as a function of wavenumber. We start by examining the solutions at high wavenumbers. The panels
d. The short-wave cutoff



We note that for the case of zero β, the short-wave cutoff corresponds to the value given by the Eady problem (in its two-layer version), namely, lo = (2 − m2π2/L2)1/2 corresponding to AT = 0. For small β, as Fig. 3 shows, the eigenfunctions are largely barotropic near the short-wave cutoff. The interesting consequence of (3.4) is that the position of the short-wave cutoff shifts to larger values of y wavenumber as β is increased. Figure 4 shows a plot of the critical value of y wavenumber, l, (on the abscissa) vs the value of β on the ordinate. In this particular setting L = 100 and m = 1. For β = 0 the solution asymptotes to the Eady value, very near 21/2 for this broad channel and then increases as β increases. The results shown here agree remarkably well with our numerical results for the cutoff. The branch point character of the critical curve can be used to obtain a splitting of the roots for smaller wavenumbers than the critical value in much the same manner as the perturbation method in Pedlosky (1970). For the sake of brevity those details are not given here but a similar analysis is given in section 4 in our discussion of the asymptotic solution for a very broad channel.
e. Longwave cutoff
We have noted that our results indicate that, as the wavenumber of the unstable wave is reduced, the phase speed of the mode tends to either the maximum or minimum value of the current, that is, either 0 or 1. At that point the mode becomes stable and the eigenfunction is limited to the layer whose current speed is not equal to cr. We exploit that fact to first find an analytical expression for the long-wave cutoff. More importantly, the analysis and its perturbation to find the imaginary part of c near the cutoff emphasizes the relationship between the unstable mode and the Rossby normal mode in the channel.
Consider now the limit in which cr → 0 at a wavenumber, lc, to be determined.







4. The broad channel approximation and the shortwave cutoff




5. Results
a. Dependence on β
Were the channel oriented in the zonal direction the flow would be stable if β were greater than unity. Figure 5 shows the dispersion curves for various values of β as well as the eigenfunction for the most unstable wave. We note that even for β = 1.5 the flow is still unstable. The growth rate is reduced for β > 1, but there is a tendency for the most unstable mode to shift to higher wavenumber and this somewhat compensates for the reduction in the value of ci in determining the growth rate. For β < 1 the most unstable mode is the gravest mode in the channel; this is no longer true for β > 1 and for each value of β it is difficult to predict a priori either the most unstable wavenumber or the character of the cross-stream structure. For the case where β = 1.5, it is clear that the most unstable mode is strongly western intensified. There are, moreover, many unstable modes in the short-wave interval for β > 1 with each mode being unstable in a small interval of y wavenumber. Figure 7 shows in detail the curves for phase speed and growth rate for β = 1.5 for 1.2 < l < 1.5 while the panels to the right and also below show the eigenfunctions at the labeled points. This interval in wavenumber contains a large number of modes. As the wavenumber is altered, branches of the phase speed coalesce and become unstable. Moving from the points marked 4 and 3 to point 2 in the upper right of the phase speed window to the points
b. Channel width
For the case of a zonal channel there is a minimum width of the channel below which the flow is stable. This is the Eady short-wave cutoff and would occur at L = π/21/2 = 2.222. When the flow is meridional and β is not zero, even a very narrow channel can still support instability for a narrow range of wavenumbers. Figure 8 shows the eigenfunctions and lists the growth rates for various channel widths. In the first panel the case where L = 0.5 is indicated. At these narrow channel widths we have found that instability occurs when two neutral modes coincide with the same frequency (or real phase speed at a given l). This is shown in Fig. 9. For the case L = 2, a channel width for which the zonally oriented flow would be stable, the range l = [0.35–0.52] is now unstable. The associated eigenfunctions are shown in Fig. 8. For very large L, for example, L = 50, the eigenfunction is similar to that at L = 10 except that the function occupies a smaller zone of the total channel in its western region.
6. Unequal layer depths


We have redone our calculations for the case where H1/H2 = 0.25, representative of the thermocline and deep ocean. The equal layer limit might be better appropriate for an attempt to resolve instabilities within the thermocline itself. Figure 10 shows the numerically calculated dispersion relation for L = 10 and β = 0.5. The growth rates are not significantly different than the equal layer case. However the along-channel wavenumber of the most unstable mode has been reduced from l = 0.99 to l = 0.8. A similar reduction is seen at the higher value of β (1.5) shown in Fig. 11. The most unstable wave now lies on a branch with phase speed >0.5 and this produces eigenfunctions that slope in the x–y plane oppositely to those previously seen, which had phase speeds equal to or less than 0.5. The eigenfunctions are again western intensified. The long-wave cutoff is not altered by the unequal layer thickness since the problem for that cutoff is independent of the parameters Fn.
7. The initial value problem
In order to verify the eigenvalue calculations, especially in the case where β > 1, we have carried out integrations of Eqs. (2.3) when the initial condition consists of a disturbance of a single l in the y direction. The initial perturbation was a sine function in x satisfying the boundary conditions. The equations were finite differenced in x (Δx = 0.1) and a second-order Runge–Kutta scheme was employed to march in time. Figure 12 shows the evolution in time of the maximum amplitude as a function of time in the panels to the left while the right-hand panels show the corresponding most unstable modes. After some time the growth is seen to approach the growth rate predicted by the normal mode theory, which is indicated by the dotted curve in the upper-left panel. In the first two panels the calculation is done for equal layer thickness. Due to the symmetry of the problem, there are two eigenfunctions sharing the same maximum growth rate but with distinct phase speeds and both emerge from the initial data. This leads to the linear vacillation of the amplitude around the predicted growth rate as the two modes alternately reinforce and interfere. On the other hand, in the last panel for the unequal layer case and for β = 1.5, there is only a single mode with the largest growth rate and the solution monotonically approaches the normal mode growth rate.
It is natural to suppose, and Spall (2000) has observed this behavior, that, if the channel is broad and the initial perturbation is also broad in the cross-stream direction compared to the deformation radius but is still confined initially to not feel the presence of the boundaries, that the initial growth that would be self-selected would be of a disturbance with very small cross-channel wavenumber. That is, locally the disturbance would look like an Eady mode and hardly sense the effect of β until the disturbance naturally broadens or propagates to interact with the boundary. Figure 13 show the results of a initial calculation in a channel 50 times wider than the deformation radius. The initial disturbance is a Gaussian in each layer, broader than the deformation radius but narrow enough to initially avoid interacting with the boundaries of the channel. The lower panel shows the growth of the disturbance at a value of l corresponding to the most unstable mode at this value of β = 0.5. We see that the initial growth matches the Eady growth rate as the disturbance first evolves with nearly no x variation within the Gaussian envelope. As time goes on and the disturbance senses the boundaries of the channel, the rate of growth approaches the normal mode growth rate that we have found from our analysis. Note the signature of the amplitude vacillation due to the presence of the superposition of the two normal modes with the same growth rate but diverse phase speeds. Figure 14 shows the evolution of the disturbance in the channel and the eventual emergence of the western intensified mode. Of course for smaller channel widths the eventual emergence of the normal mode will occur more rapidly.
8. Summary and discussion
The normal mode problem for the instability of meridional flows, even in this simple model where the flow has no horizontal shear, has a significantly different character than the zonal flow example first described by Phillips (1954). In the zonal flow case a number of a priori conditions for instability establish a threshold for the shear below which the flow must be stable. We have seen in the meridional flow case that there is no such threshold. It appears that for any value of β arbitrarily weak shears will become unstable although the associated growth rates will be small. We have not been able to prove the nonexistence of a threshold, but the combination of our failure to do so and the results of direct calculation strongly suggest its absence. We speculate that, from one point of view, this results from the destabilization of the Rossby normal modes that would exist in the channel in the absence of shear. This is further suggested by our analytical determination of the long-wave cutoff. This cutoff arises when a Rossby normal mode, limited to one layer, has a purely advective frequency (Vl) of the mean flow in one of the layers (either 0 or 1). In addition the appearance of instability for narrow channels reinforces that suggestion.
Unstable modes appear for an extended range of wavenumbers beyond the Eady cutoff when the flow is meridional and β is not zero in distinction to the zonal flow case, and again we believe this is due to a destabilization of the Rossby modes.
From another viewpoint it is worth pointing out that the potential vorticity gradients in the two layers are never colinear, so particle trajectories sensing potential vorticity gradients of opposite sign are always available. Although such opposition is necessary for zonal flows, in the absence of a stability theorem requiring such opposition in the meridional case that observation is only suggestive.
One of the fascinating features of the instability of the meridional flow is the strong dependence of the horizontal structure of the modes on the wavenumber and growth rate of the disturbance. The growing modes tend to be intensified in the western side of the domain. One can conjecture that this can be thought of as the result of the constant growth as a disturbance moves westward. The reflection of ever larger unstable disturbances from the western boundary would render the western boundary a source of perturbations larger than the arriving perturbations from the east leading to the observed western intensification. A somewhat similar result is seen in a nonmodal reflection process (Pedlosky 1993).
We have presented these results in an attempt to establish the basic properties of the normal modes for confined meridional flow. It is quite likely that for very broad channels or currents, the initial stage of growth in which the disturbance arranges itself to minimize the role of β will proceed to a stage of large enough amplitude such that secondary instabilities and nonlinear interactions will intervene before the normal modes of the present study may manifest themselves. For narrower regions or narrower currents that is less clearly the case.
We believe the subtle interactions between the nonzonal character of the flow and the β effect, illustrated by the normal modes described here, may be of general significance for the dynamics of nonzonal flows. It will be of special interest to extend the results of the present paper to include the effects of lateral structure of the basic current and the further nonlinear evolution and equilibration of the modes.
This research was supported in part by NSF OCE 9901654. The authors wish to thank Mike Spall for several valuable conversations. A portion of this work is a part of the thesis of AW in fulfillment of a Master of Science degree of the MIT/WHOI Joint Program in Physical Oceanography.
REFERENCES
Abramowitz, M., , and I. A. Stegun, 1972: Handbook of Mathematical Functions. Dover, 1046 pp.
Kamenkovich, I. V., , and J. Pedlosky, 1996: Radiating instability of nonzonal ocean currents. J. Phys. Oceanogr., 26 , 622–643.
Pedlosky, J., 1964: The stability of currents in the atmosphere and the ocean: Part I. J. Atmos. Sci., 21 , 201–219.
Pedlosky, J., . 1970: Finite amplitude baroclinic waves. J. Atmos. Sci., 27 , 15–30.
Pedlosky, J., . 1987: Geophysical Fluid Dynamics,. Chapter 7, Springer Verlag, 710 pp.
Pedlosky, J., . 1993: The reflection of unstable baroclinic waves and the production of mean coastal currents. J. Phys. Oceanogr., 23 , 2130–2135.
Phillips, N. A., 1954: Energy transformations and meridional circulations associated with simple baroclinic waves in a two-level, quasi-geostrophic model. Tellus, 6 , 273–286.
Shepherd, T. G., 1988: Nonlinear saturation of baroclinic instability. Part I: The two layer model. J. Atmos. Sci., 45 , 2014–2025.
Spall, M. A., 1994: Mechanism for low-frequency variability and salt flux in the Mediterranean salt tongue. J. Geophys. Res., 99 ((C5),) 10121–10129.
Spall, M. A., . 2000: Generation of strong mesoscale eddies by weak ocean gyres. J. Mar. Res., 58 , 97–116.
APPENDIX
The Function T in (3.12)


The eigenvalues for real c (upper) and imaginary c (lower) for the case where β = 0.5 and L = 10 (a channel width of ten deformation radii) for the case of equal layer depths as a function of the along-channel wavenumber l. Note that lci is the growth rate
Citation: Journal of Physical Oceanography 32, 3; 10.1175/1520-0485(2002)032<1075:IOMBC>2.0.CO;2

The eigenvalues for the first six unstable modes for the same parameters as in Fig. 1. Note the coalescence of the imaginary parts of the phase speed at the points where the real parts of the phase speed diverge from the value c = 0.5
Citation: Journal of Physical Oceanography 32, 3; 10.1175/1520-0485(2002)032<1075:IOMBC>2.0.CO;2

(upper left) The eigenfunctions corresponding to the eigenvalues at points indicated. Each point is shown by a double panel, the left-hand section of which is the eigenfunction in the upper layer and the right-hand side shows the panel in the lower layer. See text for a full description
Citation: Journal of Physical Oceanography 32, 3; 10.1175/1520-0485(2002)032<1075:IOMBC>2.0.CO;2

The short-wave cutoff as a function of β for a channel width L = 100. Note the increase in the cutoff wavenumber with increasing β
Citation: Journal of Physical Oceanography 32, 3; 10.1175/1520-0485(2002)032<1075:IOMBC>2.0.CO;2

The eigenvalues (left) for various values of β (0.1, 0.5, 0.9, 1.1, 1.5). (right) The most unstable eigenfunction at each value of β. In the left hand panel a box is placed at the position predicted by (3.10) in agreement with the numerical results
Citation: Journal of Physical Oceanography 32, 3; 10.1175/1520-0485(2002)032<1075:IOMBC>2.0.CO;2

Plot of cr (dashed) and ci (solid) near the long-wave cutoff for β = 0.9 and L = 10
Citation: Journal of Physical Oceanography 32, 3; 10.1175/1520-0485(2002)032<1075:IOMBC>2.0.CO;2

The detailed dispersion relation (upper left C) and eigenfunctions at selected wavenumbers for β = 1.5, L = 10
Citation: Journal of Physical Oceanography 32, 3; 10.1175/1520-0485(2002)032<1075:IOMBC>2.0.CO;2

Eigenfunctions for various channel widths (each scaled with the deformation radius). The growth rates are labeled as ω. Note that in the last case, L = 50, the panels for each layer are placed below one another
Citation: Journal of Physical Oceanography 32, 3; 10.1175/1520-0485(2002)032<1075:IOMBC>2.0.CO;2

The dispersion relation for narrow channel widths corresponding to L = 0.5 and L = 2 for both of which the zonal flow case would be stable
Citation: Journal of Physical Oceanography 32, 3; 10.1175/1520-0485(2002)032<1075:IOMBC>2.0.CO;2

The dispersion diagram for β = 0.5 for the case of unequal layer thickness, H1/H2 = 0.25, and L = 10
Citation: Journal of Physical Oceanography 32, 3; 10.1175/1520-0485(2002)032<1075:IOMBC>2.0.CO;2

As in Fig. 10 but for β = 1.5
Citation: Journal of Physical Oceanography 32, 3; 10.1175/1520-0485(2002)032<1075:IOMBC>2.0.CO;2

The results of the initial value problem. The left-hand panels show the maximum amplitude as a function of time from the calculation. The predicted growth rate from modal theory is shown as the dotted line. The middle panels show the most unstable eigenfunctions. The top row shows the case β = 0.5, l = 0.99 with predicted growth rate = 0.173. The middle row shows the case for β = 1.5 and l = 1.23, predicted growth rate of 0.05. In these two cases the layer thicknesses are equal. The bottom row show β = 1.5, l = 1.015 for the case H1/H2 = 0.25
Citation: Journal of Physical Oceanography 32, 3; 10.1175/1520-0485(2002)032<1075:IOMBC>2.0.CO;2

The results of an initial value calculation in a broad channel (L = 50). (top) The cross-stream gaussian structure of the initial perturbation in x (l = 0.955, β = 0.5). (bottom) The observed evolution with time of the disturbance. The growth rate for the Eady model with this l is 0.29 while the normal mode for the meridional channel with this value of β has a growth rate 0.193. Both are shown as dotted lines in the lower panel. Note the initial growth matches the Eady mode and then asymptotes to the modal growth rate deduced from normal mode theory
Citation: Journal of Physical Oceanography 32, 3; 10.1175/1520-0485(2002)032<1075:IOMBC>2.0.CO;2

The evolution with time of the eigenfunction described in Fig. 13
Citation: Journal of Physical Oceanography 32, 3; 10.1175/1520-0485(2002)032<1075:IOMBC>2.0.CO;2
Woods Hole Oceanographic Institution Contribution Number 10472.