In a career spanning more than 50 years, Professor Gray made extensive contributions to the study of tropical meteorology and tropical cyclones and ushered in a generation of young scientists.
Through a long career from his first paper in 1962 (Gray 1962) to his last paper in 2017 (Klotzbach et al. 2015), William M. Gray was a towering figure in tropical meteorology, particularly for global studies on tropical cyclones (TCs). A major aspect of his legacy is that he is the scientific father of a generation of scientists (Hart and Cossuth 2013), having supervised more than 70 M.S. and Ph.D. students, many of whom have become leaders in research as well as in government, university administration, and weather forecasting.
His most important legacy, however, is his personal contributions to the advancement of knowledge in tropical meteorology in general and in TC science in particular. In this paper, we discuss the scientific advancement in several key areas where Gray was a fundamental contributor: cumulus–convective-scale interactions, observational methodologies and techniques, TC inner-core structure, TC track or motion, tropical cyclogenesis, seasonal TC prediction, and climate change impact on TCs. In each case, we summarize the intellectual contribution of Gray and put it in the context of the current state of knowledge.
TROPICAL CONVECTION–LARGE-SCALE SURROUNDING CIRCULATION.
A primary area of Gray’s research in the 1970s was the interaction between cumulonimbus convection and the surrounding circulation. He made three major contributions that stimulated research and left a legacy, described in the following three subsections.
Structure of tropical weather systems.
In a series of papers, Gray and his research team documented the structure of oceanic tropical weather systems using radiosonde compositing (as described in the “Tropical cyclone composite studies” section below; Williams and Gray 1973; Gray et al. 1975; Ruprecht and Gray 1976a,b; McBride and Gray 1980). Within a scale of several hundred kilometers, the weather systems identified were cloud clusters as defined by the large areas of upper-level cloud identified through satellite imagery (Williams and Gray 1973). In recent decades, a similar system would be described as a mesoscale convective system (MCS). Modern studies focus on the structure of the embedded squall lines including downdrafts, stratiform regions, and midlevel mesoscale convective vortices [see review of Houze (2004)]. The structure documented by Gray represents the parent synoptic-scale structure, or a “grid-scale average” over the mesoscale components.
The basic dynamic structure documented was a two-layer structure, with inflow (convergence) and cyclonic vorticity from the surface up to approximately 400 hPa, capped by a layer characterized by divergence and anticyclonic vorticity. This had several implications that were revolutionary at the time, including that
the boundary layer inflow is a contributing, but not dominant, component of the mass convergence into the weather system;
the maximum vertical velocity of the system is in the mid- to upper troposphere; and
through the approximate balance between diabatic heating and vertical motion (Frank 1980; Fraedrich and McBride 1989; Sobel et al. 2001), the heating profile also has a maximum in the mid- to upper troposphere.
The implications of this two-layer structure are far reaching and implied in all subsequent dynamical models of large-scale tropical convective structure. In particular, the findings described in points 2 and 3 above influenced the development of cumulus parameterization through the requirement that the cumulus heating profile is at a maximum in the mid- to upper troposphere (Yanai et al. 1973; Arakawa and Schubert 1974; Johnson 1984; Frank and McBride 1989; Houze 1989).
The thermodynamic structure revealed in these studies is weak temperature perturbations (∼1°C) that were centered around 300 hPa. The major thermodynamic finding is that the large-scale areas containing tropical convection (the cloud clusters) are significantly more humid through the depth of the troposphere than the surrounding clear areas or regions containing no tropical convection. While seemingly trivial in modern days, this finding had major implications and still motivates modern research on convection. For example, the introduction to a major work on moisture–convection feedbacks in the tropics (Grabowski and Moncrieff 2004) cites this finding as being the basic observation motivating their theoretical development. A second example is in the developing science of aggregation of convection, where the large relative humidity differences between convective and nonconvective regions play a significant role (Tobin et al. 2012; Wing and Emanuel 2014).
In recent times, the close in situ relationship between column-integrated water vapor and the presence of tropical deep convection is so well established that it is usually implied rather than explicitly cited. Still, as stated by Muller et al. (2009, p. 1), “[a]spects of this relationship are an integral part of theories for explaining tropical phenomena including the MJO, convectively coupled waves and hurricanes.”
Documenting and understanding tropical weather systems was an area of activity across several groups at the time. Besides the work described in this subsection by Gray and collaborators at Colorado State University, parallel studies making some of the same findings were carried out at the University of California, Los Angeles (Yanai et al. 1973), and at the University of Washington (Reed and Recker 1971).
Convective mechanisms for heating and drying the large scale.
Heat and moisture budgets in the vicinities of tropical cloud clusters were determined from rawinsonde data by several investigators in the early 1970s. Gray (1973) suggested that the budgets that were obtained could be explained only if an average cloud affected its large-scale environment through a combination of dry compensating subsidence, which warmed the environment, and detrainment of hydrometeors, which evaporated and thereby cooled the environment.

Gray’s original figure showing the requirement for additional (left) upward and (right) downward subgrid-scale motion necessary to satisfy tropical heat and moisture budgets. The additional upward motion is hypothesized to take place in convective updrafts, while the downward motion includes clear air compensating subsidence, as well as moist downdrafts. From Gray (1973).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1

Gray’s original figure showing the requirement for additional (left) upward and (right) downward subgrid-scale motion necessary to satisfy tropical heat and moisture budgets. The additional upward motion is hypothesized to take place in convective updrafts, while the downward motion includes clear air compensating subsidence, as well as moist downdrafts. From Gray (1973).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
Gray’s original figure showing the requirement for additional (left) upward and (right) downward subgrid-scale motion necessary to satisfy tropical heat and moisture budgets. The additional upward motion is hypothesized to take place in convective updrafts, while the downward motion includes clear air compensating subsidence, as well as moist downdrafts. From Gray (1973).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
Forty years later, this basic mechanism is still the paradigm for how cumulonimbus clouds affect the heat and moisture balances of the surrounding troposphere (though, of course, in high-resolution simulations, the mechanisms are resolved rather than parameterized; see, for example, Plant 2010 and Arakawa and Jung 2011). The catchphrase used by Gray to his students and colleagues was that cloud impacts were carried out by a subgrid-scale up-moist, down-dry circulation. Figure 2 illustrates this mechanism for temperature balance in a skew T–logp diagram.

The “up-moist, down-dry mechanism” whereby clouds transfer heat to the surrounding circulation. The upward motion along a moist adiabat represents the vertical motion within a cumulonimbus cloud. The downward motion along a dry adiabat represents the path of the compensating between-cloud motion. Along this path, the atmosphere is warmed by adiabatic compression. The closing constant-pressure path shows the balancing cooling mechanism of radiative cooling and evaporation of detrained cloud droplets. From Gray (1973).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1

The “up-moist, down-dry mechanism” whereby clouds transfer heat to the surrounding circulation. The upward motion along a moist adiabat represents the vertical motion within a cumulonimbus cloud. The downward motion along a dry adiabat represents the path of the compensating between-cloud motion. Along this path, the atmosphere is warmed by adiabatic compression. The closing constant-pressure path shows the balancing cooling mechanism of radiative cooling and evaporation of detrained cloud droplets. From Gray (1973).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
The “up-moist, down-dry mechanism” whereby clouds transfer heat to the surrounding circulation. The upward motion along a moist adiabat represents the vertical motion within a cumulonimbus cloud. The downward motion along a dry adiabat represents the path of the compensating between-cloud motion. Along this path, the atmosphere is warmed by adiabatic compression. The closing constant-pressure path shows the balancing cooling mechanism of radiative cooling and evaporation of detrained cloud droplets. From Gray (1973).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
Diurnal variation of cumulonimbus convection and theory of convective–radiative feedbacks.
Gray and Jacobson (1977) wrote a seminal paper documenting that oceanic deep tropical convection experiences a single-cycle diurnal variation, with an early morning maximum and early evening minimum. Fundamental as this observation was, it was unknown at that time because of the masking of the signal by diurnal heating of island convection and out-of-phase contributions from stratiform and shallow convective rainfall. This paper comprehensively drew from global tropical rainfall data for small islands and corroborating information from an extensive literature review. This paper’s influence is profound, as illustrated by over 500 citations in the peer-reviewed literature.
Possibly of more lasting importance, however, is the theoretical mechanism proposed by Gray and colleagues to explain the diurnal cycle (Gray and Jacobson 1977; Fingerhut 1978; Foltz and Gray 1979; McBride and Gray 1980). They proposed that deep convection is driven by diurnally varying horizontal pressure gradients that respond to the radiative–convective heating profile differences between cloud-covered and surrounding cloud-free areas. As discussed in many recent reviews (e.g., Nesbitt and Zipser 2003; Ruppert and Johnson 2016), there is still no agreement on the physical cause of the observed diurnal variation of deep convection. However, in all reviews, the Gray and Jacobson (1977) cloud-clear radiative–convective heating differences is one of the major mechanisms evaluated. It is also invoked in ongoing studies of the diurnal cycle of TC upper-level clouds, including the phenomenon of outward-propagating diurnal pulses (Steranka et al. 1984; Kossin 2002; Dunion et al. 2014; Navarro and Hakim 2016).
The Gray–Jacobson mechanism is also important in climate change studies and associated radiative–convective feedbacks. This is discussed at length in Stephens (2005) and Stephens et al. (2008).
TROPICAL CYCLONE COMPOSITE STUDIES.
Development of the global rawinsonde network during World War II provided the first widespread atmospheric measurements of the troposphere. However, tropical rawinsonde stations were much too far apart and irregularly spaced to resolve the three-dimensional structure of individual weather systems. One answer to this problem was to composite the data. This section discusses Gray’s development of the rawinsonde compositing approach, its application to analyzing tropical cloud clusters, and its revelation of basic TC structure and dynamics.
By the 1960s, there was general agreement that latent heat release was the dominant energy source realized within the tropical atmosphere and that precipitation occurred primarily within cumulonimbus clouds. Riehl and Malkus (1958) estimated that roughly 1,500 individual thunderstorms would have to be active at all times to explain the observed mean vertical structure of the tropical atmosphere. But as the quantity and quality of satellite data increased, it quickly became obvious that deep convection in the tropics was far more organized and complex than expected. For example, the imagery showed ubiquitous mesoscale clusters of precipitating systems rather than broad fields of single-cell thunderstorms. At the time, these mesoscale areas of deep convection, typically a few hundred kilometers in diameter, were dubbed cloud clusters (see the “Tropical convection—large-scale surrounding circulation” section). Their discovery called into question the tenets of the separation of scale arguments and began a healthy, if sometimes fierce, debate on the true nature of tropical weather systems.
Gray believed firmly that while data from the operational rawinsonde network were far too sparse to resolve individual tropical weather systems, one could achieve the necessary data density by compositing data from many similar systems. This approach required enormous quantities of data and adequate computing facilities to analyze them. Gray was the first researcher to assemble a nearly complete dataset of rawinsonde data within the northwestern Pacific, southwestern Pacific, southeastern Indian, and Atlantic basins.
The first problem was data acquisition. Most rawinsonde data were stored as ink on paper at individual weather stations. Gray exerted great effort and solicited considerable funding to accomplish the tedious job of acquiring soundings and surface data from tropical weather stations and digitizing them. He, his students, and several visiting scientists then used these data to produce three-dimensional analyses of cloud clusters and TCs.
Gray’s first composite analyses were of the newly discovered tropical cloud clusters. Studies such as Williams and Gray (1973) and Ruprecht and Gray (1976a,b) used a rectangular grid centered on each of many cloud clusters (located using satellite imagery). Rawinsonde data from nearby stations were added to the gridded dataset at their positions relative to the cluster. In this manner, a dense field of data could be obtained that represented the mean, three-dimensional atmospheric state within and around the cloud cluster. This type of compositing had two obvious weaknesses. First, if essential features of the cluster circulation were randomly distributed, they would not be resolved by blending data taken from many different cloud clusters. Second, since the sampled clusters occurred at different locations, the large-scale mean, or background flow, was somewhat distorted. It was then up to the compositor to try and minimize these effects by choosing only weather systems with similar observed characteristics and from limited spatial areas using both subjective and objective criteria. Careful data synthesis elucidated clear comparisons, such as composited humidity values from tropical cloud clusters in the northwest Pacific and North Atlantic contrasted to the humidity profile of the Jordan (1958) mean sounding for the West Indies region as shown in Fig. 3 (Gray et al. 1975). There are striking differences between the humidity values at all levels above the boundary layer. These notable differences between observed air masses and the Jordan mean sounding were confirmed by Dunion and Marron (2008) when investigating Saharan air layer and non-Saharan air layer soundings.

Composited humidity values from tropical cloud clusters in the northwest Pacific and North Atlantic compared with the humidity profile of the Jordan mean sounding for the West Indies region (from Gray et al. 1975).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1

Composited humidity values from tropical cloud clusters in the northwest Pacific and North Atlantic compared with the humidity profile of the Jordan mean sounding for the West Indies region (from Gray et al. 1975).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
Composited humidity values from tropical cloud clusters in the northwest Pacific and North Atlantic compared with the humidity profile of the Jordan mean sounding for the West Indies region (from Gray et al. 1975).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
Although instrumented aircraft had probed the cores of TCs since the late 1950s, the datasets were largely confined to small areas of the cores at one or two low levels. Further, the storms could only be sampled for a few hours each day because of the logistical limitations of remotely based aircraft. Several researchers, including Gray, performed a great deal of innovative analyses of TC cores using aircraft data. However, they were unable to perform three-dimensional analyses of the storm core. The larger-scale structure was not observed, and it was impossible to compute budget analyses because of the inability to measure divergence in a vertical column.
The TC was an excellent subject for composite analysis despite the relative lack of observations over the oceans. These storms have a much more stable, slowly varying structure than do loosely organized convective systems. Further, many of the major structural features are common from one storm to the next. The positions of the circulation centers are relatively well known, and they occur during limited seasons. In recent years, various reanalysis projects have been undertaken to better assess the location and intensity of TCs around the globe (Landsea et al. 2004; Truchelut and Hart 2011; Truchelut et al. 2013).
Most of the TC composites performed by Gray and his students used a cylindrical grid extending to 15° radius (∼1,670 km) from the storm center and from the sea surface to about 80 hPa. The grid was centered on the storm circulation center at each rawinsonde observation time. In the northwest Pacific basin, even a grid this large typically had only five rawinsondes at a single time. However, composites performed for 10 years of mature typhoons in the basin could include almost 8,000 soundings. The TC composites resolved their three-dimensional structures, including the wind, moisture, and temperature fields (e.g., Frank 1977a,b). The wind measurements were accurate enough to permit computations of divergence and vorticity and hence vertical velocity. By compositing the products of the radial wind and other quantities (e.g., moisture, tangential wind, temperature), it was possible to estimate horizontal eddy fluxes. Three-dimensional budget analyses allowed computation of difficult-to-resolve quantities, such as sea surface mean fluxes.
The early composites showed that TCs were much larger than commonly thought. Their circulations clearly extended to the limits of the 15° grid boundary. This was most dramatic in the upper-tropospheric outflow layer. Later analyses compared the mean structures of developing tropical depressions to those of mature TCs. Figure 4, taken from Frank (1982), showed the radial wind differences between a mature North Atlantic hurricane and a prestorm cloud cluster in the same region. The increase in the secondary circulation as the storm spins up was confined to within about a 6° radius. At larger radii, the secondary circulation actually became somewhat weaker as the storm intensified. The same compositing technique was applied to study differences between developing and nondeveloping cloud clusters (e.g., McBride and Zehr 1981). These studies showed that while vertical wind shear directly over the circulation center of developing clusters was small, the upper-tropospheric flow exhibited strong anticyclonic shear. In contrast, clusters with weak vertical shear and weak upper-level anticyclonic shear did not develop into TCs. The compositing techniques also were used in studies of TC motion, TC structure analyses using northwest Pacific reconnaissance flight data, and TC-spawned tornados, as discussed in the “Tropical cyclone inner-core structure” and “Tropical cyclone track” sections.

Radial wind difference (in m s−1) between a mature North Atlantic hurricane and a prestorm cloud cluster in the same region (from Frank 1982).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1

Radial wind difference (in m s−1) between a mature North Atlantic hurricane and a prestorm cloud cluster in the same region (from Frank 1982).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
Radial wind difference (in m s−1) between a mature North Atlantic hurricane and a prestorm cloud cluster in the same region (from Frank 1982).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
TROPICAL CYCLONE INNER-CORE STRUCTURE.
By the early 1970s, despite an accumulation of reconnaissance flight data, structural characteristics and variability of the TC inner core remained unexamined in a systematic way. The composite philosophy also proved insightful to deciphering inner-core patterns of TCs. Three examples and their implications toward understanding TC physics and operational implementation will be discussed in this section.
Gray and Shea (1973a,b) utilized 533 radial flight legs executed by the National Hurricane Research Laboratory over a 13-yr period (1957–69) to develop composites of tangential and radial winds, D-values (difference between actual altitude above mean sea level and an altitude calibrated to a surface pressure of 1,013 hPa), and adjusted temperatures with respect to the radius of maximum wind (RMW). These were further stratified by intensity change, intensity, translation speed, and other parameters to identify useful features. Some conclusions were that
storm inflow is confined almost exclusively to the boundary layer and occurs from the RMW outwards then converges near the RMW;
the warmest TC temperatures result from subsidence and occur just inside the eyewall cloud edge where the sinking is strongest, and inner-core winds are shown to have a natural asymmetry beyond that induced by storm motion;
large vertical moist instability is present in the eyewall cloud;
supergradient winds are present at the RMW;
substantial mixing occurs between the eye and eyewall; and
inner-core heating comes from the sinking motion within the eye and not from cumulus updraft diffusion.
Gray and Shea (1973a,b) created spirited debate regarding the flight data navigation positioning accuracy and frame-of-reference gradient balance philosophies (Willoughby 1990; Gray 1991; Willoughby 1991). The composite philosophy of flight measurements encouraged future reconnaissance data to be explored in this fashion, ultimately proving an effective methodology for gleaning thermodynamic and dynamic properties of inner-core physics. In particular, the National Oceanic and Atmospheric Administration/Atlantic Oceanographic and Meteorological Laboratory/Hurricane Research Division (NOAA/AOML/HRD) flight-level data and Doppler radar archives have generated many studies on kinematic structure (Eastin et al. 2005; Aberson et al. 2006; Rogers et al. 2012). Global positioning system dropwindsondes (Hock and Franklin 1999) showed that a 0.5–1.0-km wind speed maximum is a common feature of the TC boundary layer, from which empirical adjustments for estimating 10-m winds have been developed. These datasets have spurred discussion and insight into unbalanced flow. As an example, Kepert (2001) proposed a diffusion and advection mechanism for producing the wind maximum, postulating strong inward advection of angular momentum generating the supergradient flow. Schwendike and Kepert (2008) showed that the strength of the supergradient wind flow was related to the inertial stability in an investigation of Hurricane Danielle (1988) and Hurricane Isabel (2003). Northwest Pacific TC case studies in Tse et al. (2014) and Sanger et al. (2014) showed that supergradient flow dominated the TC boundary layer inside the RMW, concluding that the unbalanced dynamics are an important component in determining radial and tangential flow maximum during a storm’s evolution.
Boundary layer composites using marine observations have also clarified the quasi-isothermal inflow of TCs (Cione et al. 2000), provided radial flux distributions (Kowaleski and Evans 2015), and clarified quadrant drag coefficient saturation (Powell et al. 2003). Such insight has resulted in adjustments to boundary layer formulations used in the Hurricane Weather Research and Forecasting Model (Gopalakrishnan et al. 2013; Trahan et al. 2016).
A second example highlights the Gray research group’s philosophy of compartmentalizing complex structures—such as the TC radial wind field—into conceptual models highlighting distinct features. Merrill (1984) postulated TC wind structure with three characteristics: intensity (maximum sustained 10-m wind), size (extent of the TC vortex from RMW to radius of gale-force wind), and strength (average wind speed of the vortex), motivated by the poor correlation of TC gale-force wind radii to intensity (Fig. 5). From this work, he proposed a metric for size called the radius of outer closed isobar (ROCI). Weatherford and Gray (1988a,b) further defined the mean tangential wind velocity within a 1°–2.5°-latitude radius from the TC center as the outer-core wind strength (OCS) and noted only weak correlations with central pressure. However, the relationship between OCS and central pressure improved when eye size was considered (Fig. 6). A high correlation was also found between OCS and radius of gale-force winds, an important parameter for the U.S. Navy and ship-routing activities, as well as a critical threshold for terminating certain evacuation operations during TC landfall events.

A conceptualized view of changes in the radial profile of tangential wind in terms of eyewall peak winds and the mean tangential wind within 1°–2.5°, defined as the OCS. From Weatherford and Gray (1988a).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1

A conceptualized view of changes in the radial profile of tangential wind in terms of eyewall peak winds and the mean tangential wind within 1°–2.5°, defined as the OCS. From Weatherford and Gray (1988a).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
A conceptualized view of changes in the radial profile of tangential wind in terms of eyewall peak winds and the mean tangential wind within 1°–2.5°, defined as the OCS. From Weatherford and Gray (1988a).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1

Mean sea level pressure (MSLP) vs OCS by eye-size class. From Weatherford and Gray (1988b).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1

Mean sea level pressure (MSLP) vs OCS by eye-size class. From Weatherford and Gray (1988b).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
Mean sea level pressure (MSLP) vs OCS by eye-size class. From Weatherford and Gray (1988b).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
The above research provided motivation that gale-force wind predictions may have skill and spurred further research into multivariate relationships for wind and pressure. Members of the Gray research project and others followed with an operational-oriented focus on empirical gale-force prediction schemes (Cocks and Gray 2002; Knaff et al. 2007; Demuth et al. 2004); analyses from infrared satellite data (Mueller et al. 2006); skill assessments of outer-core winds in numerical models (Cangialosi and Landsea 2016; Sampson and Knaff 2015); and pressure–wind relationships as a function of size, RMW, latitude, translation speed, and environmental pressure (Knaff and Zehr 2007; Courtney and Knaff 2009). This new wind–pressure relationship is being used operationally by TC warning centers around the world.
Studies of TC structure climatology have also been conducted by Gray project members (Chan and Chan 2012) and by others in the meteorology community (Kimball and Mulekar 2004; Jinnan et al. 2007). From this research has evolved a TC structure dataset known as the “extended best track” used in several studies (e.g., Mohapatra and Sharma 2015; Dolling et al. 2016). The science has evolved using outer-core structure research for improving TC vortex representation in model initialization (Tallapragada et al. 2014) and for parametric models frequently used in storm surge applications (Wood et al. 2013).
One final example is the composite study of Novlan and Gray (1974) regarding tornadoes in landfalling TCs, with more details on quadrant methodology in Novlan and Gray (1975). Before this study, smaller-sampled studies had been conducted in shorter time frames (e.g., Smith 1965). However, Novlan and Gray (1974) utilized rawinsonde and pibal datasets from 1948 to 1972 for the United States and 1950–71 for Japan, the largest scope at that time. More crucially, it identified the importance of the land interface in generating tornadoes due to increased low-level vertical shear and found that one important difference between TC-generating tornadoes and tornado-free TCs was a vertical shear of 20 m s–1 between the surface and 1.5 km. As originally postulated by Smith’s limited dataset, Novlan and Gray also confirmed tornadoes concentrated in the right-front quadrant from 100 to 400 n mi (1 n mi = 1.852 km) radially from the storm center, establishing an additional hazard of concern during landfalling events. Novlan (1973) argued that two primary reasons why tornadoes concentrated in that quadrant were that it contained the strongest vertical wind shears as well as the maximum values of low-level convergence.
The primary findings of Novlan and Gray (1974) have changed little with time, even with larger and more modern datasets (e.g., McCaul 1991). However, nuances in tornado patterns have emerged, summarized by Edwards (2012, and references cited therein). These studies have established that higher concentrations occur in the outer bands because of increased low-level vertical shear, continuous banding with supercellular structure, larger convective available potential energy values, and a more pronounced diurnal cycle than in the central region. The peak time period for TC tornadoes is just prior to landfall to 24 h after landfall because of early inland spiral bands. The most productive tornadic TCs are of hurricane strength as opposed to tropical storm strength.
Later climatological research showed that typically 10–30 EF0 or EF1 tornadoes, on the enhanced Fujita (EF) scale, will occur in the spiral bands, with an occasional EF2 or EF3, indicating that special forecast attention is necessary during landfalling events. In the United States, TC tornado prediction today is a coordinated effort between the Storm Prediction Center, the National Hurricane Center, and the local National Weather Service office staged as outlooks, watches, and warnings [see details in Edwards (2012)]. However, not all landfalling TCs produce tornadoes, and the geographic coverage varies, so forecast challenges remain, requiring further research.
TROPICAL CYCLONE TRACK.
Riehl (1954, p. 345) stated that “in the mean, tropical storms move in the direction and with the speed of the steering current, which is defined as the pressure-weighted mean flow from the surface to 300 mb over a band 8° latitude in width and centered on the storm.” Based on this statement, many schemes have since been developed to predict a TC track using the “steering flow” at different levels and over different regions around the TC (e.g., Sanders and Burpee 1968; Velden and Leslie 1991). These differences resulted from the limited cases used for each of the studies.
In an attempt to find a “universal” scheme for track prediction based on this steering concept, Gray proposed that one should apply a compositing technique to study the flow surrounding many TCs. George and Gray (1976) therefore composited 10 years of rawinsonde data over the western North Pacific for TCs with different characteristics, including direction and speed of motion, intensity, and intensity change. In all the categories, the same general conclusion was made: the 1°–7° latitude azimuthally averaged winds have the best correlations with TC motion, with the 700-hPa flow having the highest correlation for speed and that at 500 hPa for the direction of motion. This was the first time such a systematic documentation of the relationship between TC motion and its surrounding flow was made. A similar conclusion was drawn by Gray for the Atlantic using a similar methodology (Gray 1977).
Chan and Gray (1982) refined such a documentation by examining only the 5°–7° latitude azimuthally averaged winds, postulating this band average would eliminate most of the TC circulation and thus be more representative of the surrounding flow. They also composited the flow based on latitude and size and included TCs in the Southern Hemisphere. In addition to the flow at individual levels, they also examined the mean tropospheric flow between 850 and 300 hPa. In all the ocean basins, similar results to those of George and Gray (1976) were found.
A more important conclusion from both studies is that although the tropospheric flow around the TC correlates very well with TC motion, contrary to Riehl’s (1954) statement, TCs very often move at an angle to their “steering” flow, as summarized by Carr and Elsberry (1990) using the results from Chan and Gray (1982) and Holland (1984; Fig. 7). Irrespective of the characteristics of the TCs, the difference vector of TC motion minus the composite steering flow is generally nonzero and points westward and poleward. In other words, in addition to steering, some other mechanism(s) must be present to cause the TC to move. Indeed, theoretical studies starting with Rossby (1948) had proposed the Coriolis force as being a mechanism for TC movement, but no observational evidence had been shown.

(a)–(d) Difference vectors of composite TC motion minus the steering flow for various categories of TC characteristics in the western North Pacific based on the data from Chan and Gray (1982): (a) latitude [north (GT) and south (LT) of 20°N], (b) direction of TC motion, (c) speed of TC motion [fast (F), medium (M), and slow (SL)], and (d) TC intensity [very intense (VI), intense (I), weak (WK)]. (e),(f) As in (a)–(d), but for TCs in the Southern Hemisphere based on the data from Holland (1984): (e) direction of TC motion, and (f) before (B), no (N), or after (A) recurvature (from Carr and Elsberry 1990).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1

(a)–(d) Difference vectors of composite TC motion minus the steering flow for various categories of TC characteristics in the western North Pacific based on the data from Chan and Gray (1982): (a) latitude [north (GT) and south (LT) of 20°N], (b) direction of TC motion, (c) speed of TC motion [fast (F), medium (M), and slow (SL)], and (d) TC intensity [very intense (VI), intense (I), weak (WK)]. (e),(f) As in (a)–(d), but for TCs in the Southern Hemisphere based on the data from Holland (1984): (e) direction of TC motion, and (f) before (B), no (N), or after (A) recurvature (from Carr and Elsberry 1990).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
(a)–(d) Difference vectors of composite TC motion minus the steering flow for various categories of TC characteristics in the western North Pacific based on the data from Chan and Gray (1982): (a) latitude [north (GT) and south (LT) of 20°N], (b) direction of TC motion, (c) speed of TC motion [fast (F), medium (M), and slow (SL)], and (d) TC intensity [very intense (VI), intense (I), weak (WK)]. (e),(f) As in (a)–(d), but for TCs in the Southern Hemisphere based on the data from Holland (1984): (e) direction of TC motion, and (f) before (B), no (N), or after (A) recurvature (from Carr and Elsberry 1990).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
Chan (1984) for the first time composited rawinsonde observations to demonstrate that the vorticity budget can be used to diagnose TC movement. A TC tends to move toward the area of maximum relative vorticity tendency, which within most of the troposphere (850–300 hPa) is mainly contributed by the advection of absolute vorticity. The advection of relative vorticity is essentially steering. The advection of planetary vorticity is the component that was not considered. It is apparently this component that causes the TC to move differently from that of steering.
Holland (1983) showed that in the Northern Hemisphere, advection of planetary vorticity by the tangential winds of the TC gives a positive (negative) vorticity tendency to the west (east) of the TC. This sets up a secondary circulation with cyclonic (anticyclonic) flow, which then drives the TC northward. Combining this movement with the relative vorticity tendency being positive to the west causes the TC to move toward the northwest. Chan (1982) discussed this concept to explain the results of George and Gray (1976) and Chan and Gray (1982) in terms of the deviation of TC motion from the steering flow.
Chan and Williams (1987) ran a barotropic numerical model to study the concept proposed by Holland (1983) further. They found that inclusion of the nonlinear advection of relative vorticity was crucial in causing the TC to move toward the northwest. This northwestward movement of a TC caused by the variation of the Coriolis parameter has been known as the beta effect. Fiorino and Elsberry (1989) showed that the asymmetric flow associated with the beta effect is in the form of a pair of counter-rotating gyres, which have since been known as “beta gyres” (Fig. 8).

Asymmetric streamfunction (104 m2 s−1) associated with the beta effect in a barotropic model. Solid and dashed contours indicate positive and negative values of streamfunction, respectively (from Fiorino and Elsberry 1989).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1

Asymmetric streamfunction (104 m2 s−1) associated with the beta effect in a barotropic model. Solid and dashed contours indicate positive and negative values of streamfunction, respectively (from Fiorino and Elsberry 1989).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
Asymmetric streamfunction (104 m2 s−1) associated with the beta effect in a barotropic model. Solid and dashed contours indicate positive and negative values of streamfunction, respectively (from Fiorino and Elsberry 1989).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
This new concept that TC motion equals steering plus the beta effect, together with their nonlinear interaction, proposed by Gray and his students triggered many subsequent theoretical and observational studies [see review by Chan (2005)]. These concepts have also been used to explain why numerical weather prediction (NWP) models for TC track prediction at the time all tended to have a slow bias in the first 24–36 h. A symmetric vortex was generally inserted into the model, but because of the beta effect, the vortex should be asymmetric. Carr and Elsberry (1992) used a barotropic model to obtain improvements in track forecasting by explicitly including the beta propagation. Following this idea, Heming et al. (1995) inserted an asymmetric vortex in the Met Office operational forecast model to simulate the beta effect and reduced the TC track forecast errors for all time periods by up to 50%. Since then, many operational NWP models for TC track predictions have also included asymmetric vortices in their initial conditions, until the last decade, when many more observations have become available to be assimilated into the model so that the initial vortex could be better represented, especially with the improved assimilation techniques.
With advances in computational capability, TC motion studies evolved from barotropic to fully baroclinic models, and the use of vorticity budgets to diagnose TC motion were replaced by potential vorticity budgets to take into account the diabatic heating in a baroclinic atmosphere (Wu and Emanuel 1995a,b; Wu and Wang 2000; Chan et al. 2002). With these studies, it may be concluded that the physical mechanisms responsible for TC motion over the ocean are largely understood (Chan 2005).
Thus, the pioneering effort of Gray and his students in initiating a composite study to relate TC motion to its environmental flow has led to a relatively complete understanding of TC motion and contributed to a substantial improvement in operational forecasting of TC tracks for close to two decades.
TROPICAL CYCLOGENESIS.
Gray’s research often focused upon understanding the physical mechanisms and controls of tropical cyclogenesis: the formation of a coherent convectively forced, synoptic-scale vortex. Gray wrote a seminal paper on tropical cyclogenesis (Gray 1968), which today is still one of the most widely cited papers in all of tropical meteorology. This foundational paper explained the observed geographic and annual cycle climatology of tropical cyclogenesis with the key environmental factors that determine these distributions. One of the most important factors was tropospheric vertical shear. Previously, the importance of vertical shear for genesis (and intensification) was not fully appreciated in either the peer-reviewed literature or by operational forecasters, though earlier work (e.g., Riehl 1954; Palmen 1956; Ramage 1959) did make some linkages. The paper also provided a clear differentiation between basins in which the monsoon trough (termed “doldrum equatorial trough” in the paper) is present and helped promote genesis versus basins in which the intertropical convergence zone (termed “trade winds equatorial trough” in the paper) dominates and is less conducive to tropical cyclogenesis. Gray further elucidated the necessary factors for genesis in Gray (1979), where he formulated the occurrence of tropical cyclogenesis to be a combination of dynamic and thermodynamic potentials (Fig. 9). Here is where he first specifically related genesis to the following factors:
cyclonic low-level vorticity (from a preexisting easterly wave, the monsoon trough, or an upper-level low/frontal boundary),
moist midtroposphere,
conditional instability through a deep tropospheric layer,
warm and deep oceanic mixed layer,
weak tropospheric vertical shear of the horizontal wind, and
location of disturbance a few degrees poleward of the equator (i.e., a significant value of Coriolis force).

Portrayal of (a) four-season sum of the original genesis parameter and (b) TC origin locations for a 20-yr climatology from 1958 to 1977. Both figures taken from Gray (1979).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1

Portrayal of (a) four-season sum of the original genesis parameter and (b) TC origin locations for a 20-yr climatology from 1958 to 1977. Both figures taken from Gray (1979).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
Portrayal of (a) four-season sum of the original genesis parameter and (b) TC origin locations for a 20-yr climatology from 1958 to 1977. Both figures taken from Gray (1979).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
These six “ingredients” in the TC “recipe” for genesis are still widely accepted today, though modified some. For example, newer formulations use subsets of these factors and obtain more variability explained, while others have combined factors together (such as 1 and 6 together as absolute vorticity) or have replaced ingredients with a more physically based factor (such as substituting in potential intensity for 4; Emanuel and Nolan 2004; Camargo et al. 2007c).
In the 1980s and early 1990s, Gray’s contributions to TC genesis were in conjunction with many of his graduate students at Colorado State University: McBride and Gray (1980), McBride (1981a,b), McBride and Zehr (1981), Frank (1982), Love (1985), Lee (1989a,b), and Zehr (1992). Much of this work demonstrated that while the thermodynamic components in the tropical cyclogenesis recipe (2, 3, and 4) were often satisfied for much of the cyclone season, it was the variability in the dynamical factors (1 and 5) that determined whether a disturbance would undergo genesis or not. Such differentiation allowed for better understanding of the physical causes between the very active TC basins (such as the northwest and northeast Pacific), the “marginal” basins (such as the North Atlantic), and basins that are generally free of TC activity (such as the southeast Pacific and South Atlantic). Zehr’s work in particular identified a two-stage process (Fig. 10) for genesis when the synoptic conditions were fulfilled. Stage one is when a convective maximum occurs within the circulation of a synoptic-scale tropical disturbance. This initiates a midlevel mesoscale vortex. Stage two occurs one day to a few days later with a second convective burst, which then develops the mesoscale vortex to the surface and thus genesis. Gray’s last direct contribution to the field (Gray 1998) was an overview of Gray and his students’ 30 years of research toward understanding the physics of genesis.

Conceptual model of tropical cyclogenesis with illustrations and descriptions of characteristics that are observable using satellite imagery (from Zehr 1992).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1

Conceptual model of tropical cyclogenesis with illustrations and descriptions of characteristics that are observable using satellite imagery (from Zehr 1992).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
Conceptual model of tropical cyclogenesis with illustrations and descriptions of characteristics that are observable using satellite imagery (from Zehr 1992).
Citation: Bulletin of the American Meteorological Society 98, 11; 10.1175/BAMS-D-16-0116.1
Despite these accomplishments, as Frank (1987, p. 78) described, “[a]ttempts to apply these criteria to daily forecasting of cyclone genesis have met [at that time] with little success.” It was not until the advent of high-resolution global weather forecast models in the late 1990s that predictions of genesis were possible. The first methods with some forecast skill were those that used the synoptic-scale output of the global models to make genesis predictions (e.g., Hennon and Hobgood 2003). More recent work is in using the explicit genesis predictions from global models after accounting for various models’ biases either in being over- or undercyclogenetic (Halperin et al. 2013). These improved capabilities have allowed operational TC forecasting centers such as the National Hurricane Center2 to begin issuing quantitative, probabilistic-based genesis predictions—48-h graphical tropical weather outlooks (GTWO) beginning in 2008 and 5-day GTWO starting in 2013 (Fig. 11). The success of these genesis predictions would not be possible without the foundation for understanding tropical cyclogenesis that Gray and his students provided over the last few decades.
