1. Introduction
Terrestrial vegetation is an important component of the climate system: vegetation alters the physical properties of the land surface, affecting how much solar radiation is received at the surface as well as how that energy is partitioned. Further, forests provide an important sink for anthropogenic carbon dioxide (CO2) emissions in part because of the fertilization effect in which higher levels of atmospheric CO2 increase photosynthesis and water use efficiency by plants (e.g., Notaro et al. 2007; Ballantyne et al. 2012). Despite these potential beneficial effects of increasing CO2, there is recent evidence of increasing tree mortality around the world in numerous ecosystem types (Allen et al. 2010). These widespread observations of mortality events suggest that climate change is contributing to increasing forest death, which may eventually lead to a tipping point where climate changes rapidly or substantially enough to initiate large-scale die-offs. Warming could accelerate through the reduction of the biomass carbon sink.
As a result of the interplay between the two effects of CO2, the fertilization effect and climate change itself, it is unclear how future climate and vegetation trajectories will unfold. In the tropics, the relative importance of two competing climate responses to increasing greenhouse gases1—a local (“bottom up”) response and a remote (“top down”) response (Giannini 2010; Seth et al. 2011, 2013)—is likely an important arbiter of how climate will change, with important implications for how vegetation will respond. In the local response, increases in net surface radiation lead to increases in evaporation and low-level moist static energy, driving increased precipitation and moisture convergence. In the remote response, the rising heat content of the ocean and higher sea surface temperatures (SSTs) result in nearly uniform warming in the upper troposphere (Sobel et al. 2001), increasing stability. Precipitation is reduced over tropical land areas where there is insufficient moist static energy to overcome this enhanced stability (Chou and Neelin 2004; Chou et al. 2009; Giannini 2010). Subsequent reductions in precipitation recycling promote further drying. The remote mechanism can operate in space, with precipitation decreasing on the subtropical margins (Chou and Neelin 2004; Chou et al. 2009) but also in time with precipitation decreasing in the winter or dry season into the spring (Biasutti and Sobel 2009; Seth et al. 2010, 2011, 2013).
These mechanisms can also operate over the oceans via the spatial pattern of SST warming; this is known as the “warm get wetter” pattern (Xie et al. 2010). Areas where SSTs increase more relative to the mean tropical warming regions can more easily sustain convection despite warming of the free troposphere. These regions with greater warming have greater instability and thus become wetter in future climate projections (Xie et al. 2010). Therefore, in this warm get wetter mechanism, the pattern of SST warming controls where precipitation will increase the most (Rauscher et al. 2008, 2011; Xie et al. 2010; Ma and Xie 2013). Additionally, these precipitation changes may affect other regions remotely through teleconnections (Huang et al. 2013).
The influence of SST warming patterns is evident in modeling studies of climate and vegetation change, with perhaps the best example being simulated die-off over the Amazon basin. In a widely cited study, Cox et al. (2000) used the Hadley Centre Coupled Atmosphere–Ocean Model version 3 (HadCM3) coupled to an ocean carbon cycle model and a dynamic vegetation model to simulate future climate and ecosystem change over the Amazon. In their simulation, a complete die-off of the Amazon forest begins in the middle of the twenty-first century. This simulated die-off was the result of severe droughts due to an El Niño–like warming over the central-eastern Pacific that altered the atmospheric circulation such that large-scale descent and a reduction in precipitation occurred over the Amazon (Cox et al. 2000, 2004; Li et al. 2006). HadCM3 also simulated a meridional gradient in the warming of the tropical Atlantic that contributed to drying (Harris et al. 2008).
However, all model simulations of future climate do not display this same SST warming pattern, and not all models simulate drying over the Amazon (Li et al. 2006). In fact, of all the models in the Coupled Model Intercomparison Project phase 3 (CMIP3; Meehl et al. 2007), HadCM3 simulates the largest future decrease in precipitation over the Amazon (Li et al. 2006). Therefore, we must ask, how probable and how uncertain are these HadCM3 results over tropical South America? More generally, how probable is large-scale vegetation change in the tropics as a whole? Since none of the scenario simulations in the CMIP3 database (and only a few in CMIP5) were run with fully interactive carbon cycles or dynamic vegetation, a large fully interactive carbon–climate multimodel ensemble is not possible.
Here we create projections of future vegetation and climate change over the tropics using a single model, the National Science Foundation (NSF)–DOE Community Earth System Model (CESM1.0; Gent et al. 2011). We mimic a multimodel atmosphere–vegetation ensemble through the use of SST projections from eight different coupled climate models in CMIP3 (Meehl et al. 2007) as boundary conditions. In this way, we can produce multiple realizations of twenty-first-century climate under a medium–high emissions scenario (SRES A2; IPCC 2000) using the interactive land and atmosphere components of CESM. Since previous studies have reported the potential for the die-off of the Amazon, we focus our efforts on tropical South America while also considering the tropics as a whole. In contrast to recent studies that have utilized offline simulations driven with bias-corrected data from CMIP3 models (e.g., Huntingford et al. 2013), we ran our simulations with full interaction between the land and atmosphere. Because we are using a single model, we focus our analysis on the effects of the differences in the climate drivers rather than on the effects of physiological parameterizations. Importantly, we examine the ability of our single-model ensemble to capture the spread of the original CMIP3 multimodel ensemble.
Section 2 describes the models and experimental design. Section 3 presents an analysis of the control simulation. The ability of our single model ensemble to capture the range of responses in the full CMIP3 ensemble is discussed in section 4. The simulated changes in tropical vegetation are discussed in section 4b. Finally, discussion and conclusions are presented in section 5.
2. Model, data, and experimental design
a. Model
For our simulations we use the atmosphere and land components of the NSF–DOE Community Earth System Model (CESM) (Gent et al. 2011), the most recent version of the Community Climate System Model (CCSM). The atmospheric component of CESM is the Community Atmosphere Model (CAM); we use the finite volume (FV) dynamical core and CAM4 physics with 26 vertical levels (Neale et al. 2010). The spatial resolution of the model is 1.9° latitude by 2.5° longitude. This medium horizontal resolution was chosen because of the need to perform 16 ensemble members, including 8 members with dynamic vegetation. Exchanges of heat, moisture, and momentum fluxes between the land and the atmosphere are simulated by the Community Land Model (CLM), version 4.0. In CLM, vegetation coverage is described in each grid cell by fractional areas of plant functional types (PFTs). There are 17 PFTs, including bare ground, 11 tree PFTs, 3 grass PFTs, and 2 crop PFTs; however, crop types and historical transient land-use change are not used when the dynamic vegetation model is enabled (Lawrence et al. 2011). CLM includes a carbon–nitrogen (CN) biogeochemical model that controls carbon and nitrogen dynamics (Thornton et al. 2007) and predicts vegetation, litter, soil carbon and nitrogen states, and vegetation phenology. We also utilize the dynamic vegetation model (CNDV), which is based on the Lund–Potsdam–Jena (LPJ) model (Sitch et al. 2003). In our simulations, the interaction between the land and atmosphere components is two way, unlike previous studies performed to evaluate the basic performance of CLM in CESM (e.g., Gotangco Castillo et al. 2012).
We performed two ensembles: one ensemble with CAM and CLM with CN and the dynamic vegetation model enabled (CAM–CLM-CNDV) (Levis et al. 2004; Gotangco Castillo et al. 2012) and one ensemble with no dynamic vegetation (CAM–CLM-CN). Carbon cycle dynamics are controlled by CAM–CLM-CN in both sets of simulations. The main differences between CAM–CLM-CN and CAM–CLM-CNDV are the biogeography as represented by the distribution of PFTs and the parameterization of mortality processes (Gotangco Castillo et al. 2012). For CAM–CLM-CN, vegetation PFTs are prescribed from satellite data and cannot change through time. In CAM–CLM-CNDV, PFTs are established through time, and vegetation change is represented by a change in the fractional PFT coverage of a grid cell at the end of each simulation year (Sitch et al. 2003). PFT types can change in CAM–CLM-CNDV because of 20-yr climate envelopes based on temperature and precipitation limits as well as because of mortality mechanisms. These mortality mechanisms differ between CAM–CLM-CN and CAM–CLM-CNDV: CAM–CLM-CNDV calculates mortality due to heat stress (based on an accumulation of growing degree days), fire, and growth efficiency, while CAM–CLM-CN uses a flat 2% mortality rate. While PFT type cannot change in CAM–CLM-CN simulations, vegetation characteristics such as leaf area index can still change because of climate change or interannual variability.
Because the fire module in CLM is important to the results discussed in section 4b, it is described in more detail here. In both CAM–CLM-CN and CAM–CLM-CNDV, fire is a prognostic algorithm based on Thonicke et al.’s (2001) intermediate-complexity Global Fire Model (Glob-FIRM), which depends on surface fuel availability and near-surface soil moisture conditions (Oleson et al. 2010); however, ignition source availability and the effects of wind speed on fire are not considered (Li et al. 2012). As with the PFT changes, the effects of fire are applied at the end of each simulation year to each grid cell. The effect of fire depends on fire season length in days, which is dependent on subhourly fire probability. This in turn is a function of the ratio of volumetric soil moisture in the top 0.5 m of soil water to the saturation soil water. Fire season length and fuel control the area affected by fire. A difference between CAM–CLM-CN and CAM–CLM-CNDV is that with CNDV fire can affect PFT coverage by removing a number of individuals as a function of the area burned from the PFT population. If enough individuals are removed, PFT cover is reduced to zero (i.e., the land cover changes to bare ground).
b. Data
To evaluate the control simulation, we use gridded land-only observations from the University of Delaware dataset, version 3.02 (Willmott and Matsuura 1995, 2001). Monthly 2-m temperature data are available at 0.5° resolution for the 1900–2010 timeframe; here we use an average from 1970 to 1999. For evaluation of precipitation, we use the Climate Prediction Center Merged Analysis of Precipitation (CMAP; Xie and Arkin 1996) dataset and the Climate Anomaly Monitoring System (CAMS) and OLR Precipitation Index (OPI; Janowiak and Xie 1999). The CMAP dataset is a blended product of global satellite and gauge data on a 2.5° × 2.5° latitude–longitude grid, approximately the same as the CAM–CLM-CN simulations. We use the period 1979–2008, since the dataset is available from 1979 onward. CAMS–OPI is a precipitation estimation technique that produces real-time monthly analyses of global precipitation. Rain gauge observations (CAMS data) are combined with precipitation estimates from a satellite algorithm (OPI). The analyses are on a 2.5° × 2.5° latitude–longitude grid, are updated each month, and are available for 1979–present.
c. Experiment design
As discussed above, recent numerical experiments suggest that future SST patterns may play an important role in shaping how precipitation may change in a warmer world (Rauscher et al. 2008, 2011; Xie et al. 2010; Ma and Xie 2013). For example, simulations that show warming in the eastern tropical Pacific and a meridional gradient in warming in the tropical Atlantic tend to reduce precipitation over the Amazon. Models that do not have a similar warming pattern do not simulate reductions in precipitation over the Amazon (Li et al. 2006). To simulate a range of potential future climates while using a single model, we used SST projections from eight different fully coupled climate models from CMIP3 (Meehl et al. 2007) as boundary conditions for our simulations.
The spinup procedures used for the simulation are described in full by Jiang et al. (2013); we summarize here for brevity. We performed a 155-yr coupled atmosphere–land (CAM–CLM-CNDV) spinup simulation. Initial conditions for this 155-yr coupled simulation were provided by a 200-yr offline CLM-CNDV simulation that cycled through the 1948–2004 observed atmospheric forcing (Qian et al. 2006) and started from the end of a twentieth-century CLM-CN transient simulation. Net ecosystem exchange (NEE) was near zero by the end of the twentieth-century control simulation.
Two ensembles were performed, each with one historical control simulation (1900–2005) and eight future scenario simulations (2005–99), as illustrated in Fig. 1. The historical control simulation in each ensemble is forced by observed SSTs with the land model initial conditions taken from the end of the spinup run. The results from the end of the single historical simulation were used to initialize the eight future scenario simulations in which SST projections from different CMIP3 models were bias corrected based on observed SSTs (Hurrell et al. 2008). These eight GCMs include the NCAR CCSM3, CNRM-CM3, MPI ECHAM5, GFDL CM2.1, GISS-ER, UKMO HADCM3, UKMO HadGEM1, and MRI-CGCM2.3.2a as shown in Fig. 1 and Fig. 2. Selection of the different models is described in Jiang et al. (2013). Throughout the text we refer to these CMIP3 simulations as the “parent” CMIP3 simulations.



Bias-corrected annual average SST anomalies (°C) for the end of the twenty-first century (2070–99) vs the end of the twentieth century (1970–99) from CMIP3 simulations (a) CCSM3, (b) CNRM-CM3, (c) ECHAM5, (d) GFDL, (e) GISS-ER, (f) HadCM3, (g) HadGEM1, and (h) MRI.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Bias-corrected annual average SST anomalies (°C) for the end of the twenty-first century (2070–99) vs the end of the twentieth century (1970–99) from CMIP3 simulations (a) CCSM3, (b) CNRM-CM3, (c) ECHAM5, (d) GFDL, (e) GISS-ER, (f) HadCM3, (g) HadGEM1, and (h) MRI.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Bias-corrected annual average SST anomalies (°C) for the end of the twenty-first century (2070–99) vs the end of the twentieth century (1970–99) from CMIP3 simulations (a) CCSM3, (b) CNRM-CM3, (c) ECHAM5, (d) GFDL, (e) GISS-ER, (f) HadCM3, (g) HadGEM1, and (h) MRI.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
The two ensembles of one control simulation and eight scenario simulations differ in the use of the dynamic vegetation model. For the first ensemble, we used CAM-CLM coupled with dynamic vegetation enabled. In the second ensemble, the dynamic vegetation was turned off. To be consistent with the future SST projections, greenhouse gas emissions from the IPCC SRES A2 emissions scenario are used as forcing (IPCC 2000). Aerosol concentrations and deposition rates in all simulations were held constant at year-2000 levels. Prescribed transient CO2 and nitrogen deposition rates (Lamarque et al. 2010) were employed for the historical and future-year simulations. Unless otherwise noted, averages representing the late twenty-first century (2070–99) and the twentieth century (1970–99) are compared to assess the future changes relative to the present.
3. Results: Control simulations
We first examine the vegetation distribution simulated in our historical control simulation with dynamic vegetation on. Overall, our control simulation results are very similar to the results obtained in a previous evaluation of CLM-CNDV by Gotangco Castillo et al. (2012, their Fig. 2). The main difference between the simulations of Gotangco Castillo et al. (2012) and those presented here is that the former utilized a one-way forcing of CLM-CNDV by atmospheric data produced using CAM, whereas our simulations are interactive between the atmosphere and land surface.
Figures 3a–f show the observed distribution of PFTs derived from MODIS satellite data that are used within the model when prescribing “natural” vegetation cover (i.e., the vegetation distribution that would be used in the CAM–CLM-CN configuration when the dynamic vegetation is turned off). Figures 4a–f show the PFTs produced in the CAM–CLM-CNDV control simulation. The spatial distribution of broadleaf evergreen trees agrees fairly well with the satellite-derived PFTs over tropical regions; that is, broadleaf evergreens are found over central America and the Amazon, central Africa, southern Asia, Southeast Asia, and parts of Australia and New Zealand. Relative fractional coverage is underestimated, however. This deficiency is compensated for by increased deciduous tree coverage. For example, while the Amazon is covered by more than 90% broadleaf evergreen according to the satellite data, in CAM–CLM-CNDV it has approximately 45% broadleaf evergreen and 45% deciduous trees. In CAM–CLM-CNDV there is no mandatory senescence period, so the deciduous trees are able to “compete” in this region (Gotangco Castillo et al. 2012). Grass (C3 and C4), shrub, and bare-ground PFTs are shown in Figs. 3d,e and Figs. 4d,e. Over the Southern Hemisphere the simulation agrees more closely with observations, with shrub cover found over southern South America, southern Africa, and Australia. Grasses are underestimated almost everywhere, whereas bare ground is overestimated. Again, these simulations compare well with the established performance of CNDV and its parent DGVM, LPJ (Notaro et al. 2007).

Observed plant function types (% cover) used in CAM–CLM-CN simulations: (a) broadleaf evergreen trees, (b) needleleaf evergreen trees, (c) deciduous trees, (d) shrubs, (e) grasses, and (f) bare ground.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Observed plant function types (% cover) used in CAM–CLM-CN simulations: (a) broadleaf evergreen trees, (b) needleleaf evergreen trees, (c) deciduous trees, (d) shrubs, (e) grasses, and (f) bare ground.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Observed plant function types (% cover) used in CAM–CLM-CN simulations: (a) broadleaf evergreen trees, (b) needleleaf evergreen trees, (c) deciduous trees, (d) shrubs, (e) grasses, and (f) bare ground.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Simulated plant functional types (average over 1970–99; % cover) in CAM–CLM-CNDV simulations: (a) broadleaf evergreen trees, (b) needleleaf evergreen trees, (c) deciduous trees, (d) shrubs, (e) grasses, and (f) bare ground.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Simulated plant functional types (average over 1970–99; % cover) in CAM–CLM-CNDV simulations: (a) broadleaf evergreen trees, (b) needleleaf evergreen trees, (c) deciduous trees, (d) shrubs, (e) grasses, and (f) bare ground.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Simulated plant functional types (average over 1970–99; % cover) in CAM–CLM-CNDV simulations: (a) broadleaf evergreen trees, (b) needleleaf evergreen trees, (c) deciduous trees, (d) shrubs, (e) grasses, and (f) bare ground.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
We can compare this CAM–CLM-CNDV simulation with our control CAM–CLM-CN simulation to determine the effect of these differences in vegetation cover (dynamic versus prescribed, respectively) on the simulation of twentieth-century climate (Figs. 5 and 6). Comparing averages for model years 1970–99, there are no statistically significant differences in temperature over most of the tropics. Precipitation differences between the two simulations are small, less than 0.5 mm day−1. Although the Amazon tree cover is spuriously split between broadleaf evergreen and deciduous tree types in the CAM–CLM-CNDV simulation as noted above, only a small region over eastern Brazil shows lower precipitation in CAM–CLM-CNDV compared to CAM–CLM-CN.

Modeled and observed annual average temperature (1970–99; °C) for (a) CAM–CLM-CNDV control simulation, (b) CAM–CLM-CN control simulation, (c) WM observations, and differences (d) CAM–CLM-CNDV − CAM–CLM-CN, (e) CAM–CLM-CNDV − WM, and (f) CAM–CLM-CN − WM. For (d)–(f), only significant differences are shown.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Modeled and observed annual average temperature (1970–99; °C) for (a) CAM–CLM-CNDV control simulation, (b) CAM–CLM-CN control simulation, (c) WM observations, and differences (d) CAM–CLM-CNDV − CAM–CLM-CN, (e) CAM–CLM-CNDV − WM, and (f) CAM–CLM-CN − WM. For (d)–(f), only significant differences are shown.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Modeled and observed annual average temperature (1970–99; °C) for (a) CAM–CLM-CNDV control simulation, (b) CAM–CLM-CN control simulation, (c) WM observations, and differences (d) CAM–CLM-CNDV − CAM–CLM-CN, (e) CAM–CLM-CNDV − WM, and (f) CAM–CLM-CN − WM. For (d)–(f), only significant differences are shown.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Modeled and observed annual average precipitation (1970–99; mm day−1) for (a) CAM–CLM-CNDV control simulation, (b) CAM–CLM-CN control simulation, (c) CMAP observations, and differences (d) CAM–CLM-CNDV − CAM–CLM-CN, (e) CAM–CLM-CNDV − CMAP, and (f) CAM–CLM-CN − CMAP. For (d)–(f), only significant differences are shown.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Modeled and observed annual average precipitation (1970–99; mm day−1) for (a) CAM–CLM-CNDV control simulation, (b) CAM–CLM-CN control simulation, (c) CMAP observations, and differences (d) CAM–CLM-CNDV − CAM–CLM-CN, (e) CAM–CLM-CNDV − CMAP, and (f) CAM–CLM-CN − CMAP. For (d)–(f), only significant differences are shown.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Modeled and observed annual average precipitation (1970–99; mm day−1) for (a) CAM–CLM-CNDV control simulation, (b) CAM–CLM-CN control simulation, (c) CMAP observations, and differences (d) CAM–CLM-CNDV − CAM–CLM-CN, (e) CAM–CLM-CNDV − CMAP, and (f) CAM–CLM-CN − CMAP. For (d)–(f), only significant differences are shown.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
4. Results: Future projections
a. Reproducibility of CMIP3 ensemble
As shown in Fig. 2, the long-term annual average of the SST change patterns used to drive the CAM–CLM-CN and CAM–CLM-CNDV simulations varies between the CMIP3 models. Some models show El Niño–like warming in the tropical eastern Pacific (e.g., GFDL, HadCM3, CNRM) and others do not (GISS-ER, ECHAM5). To see how our simulations responded to these eight different SST warming patterns compared to their parent CMIP3 simulations, we compare the precipitation changes between the end of the twenty-first century (2070–99) to the end of the twentieth century (1970–99) in our simulations with the parent CMIP3 simulations. Figure 7 shows the percent precipitation differences for the CAM–CLM-CN simulations for comparison with the parent CMIP3 simulations (Fig. 8). There are general similarities between the CAM–CLM-CN simulations and their parent CMIP3 simulations that are typical of almost all future climate change simulations. For example, drying is found in the subtropics, and the midlatitude storm tracks shift poleward. Over the oceans, the precipitation changes tend to follow departures from the zonal mean SST warming (Ma and Xie 2013). Where SSTs warm more than the zonal mean, precipitation increases, and where they warm less, precipitation decreases. These differences in warming pattern can be large and account for a substantial portion (up to one-third) of intermodel spread in future precipitation projections (Ma and Xie 2013). For example, the tropical North Atlantic shows a warming minimum in most CMIP3 projections of future SSTs (Fig. 2; CCSM3 and HadCM3 show this clearly) (Leloup and Clement 2009), which is associated with lower projected precipitation in the region (Rauscher et al. 2011).

Annual average precipitation change (%) for end of the twenty-first century (2070–99) vs end of the twentieth century (1970–99) for CAM–CLM-CN simulations driven by SSTs from (a) CCSM3, (b) CNRM-CM3, (c) ECHAM5, (d) GFDL, (e) GISS-ER, (f) HadCM3, (g) HadGEM1, and (h) MRI.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Annual average precipitation change (%) for end of the twenty-first century (2070–99) vs end of the twentieth century (1970–99) for CAM–CLM-CN simulations driven by SSTs from (a) CCSM3, (b) CNRM-CM3, (c) ECHAM5, (d) GFDL, (e) GISS-ER, (f) HadCM3, (g) HadGEM1, and (h) MRI.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Annual average precipitation change (%) for end of the twenty-first century (2070–99) vs end of the twentieth century (1970–99) for CAM–CLM-CN simulations driven by SSTs from (a) CCSM3, (b) CNRM-CM3, (c) ECHAM5, (d) GFDL, (e) GISS-ER, (f) HadCM3, (g) HadGEM1, and (h) MRI.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Annual average precipitation change (%) for end of the twenty-first century (2070–99) vs end of the twentieth century (1970–99) for CMIP3 simulations (a) CCSM3, (b) CNRM-CM3, (c) ECHAM5, (d) GFDL, (e) GISS-ER, (f) HadCM3, (g) HadGEM1, and (h) MRI.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Annual average precipitation change (%) for end of the twenty-first century (2070–99) vs end of the twentieth century (1970–99) for CMIP3 simulations (a) CCSM3, (b) CNRM-CM3, (c) ECHAM5, (d) GFDL, (e) GISS-ER, (f) HadCM3, (g) HadGEM1, and (h) MRI.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Annual average precipitation change (%) for end of the twenty-first century (2070–99) vs end of the twentieth century (1970–99) for CMIP3 simulations (a) CCSM3, (b) CNRM-CM3, (c) ECHAM5, (d) GFDL, (e) GISS-ER, (f) HadCM3, (g) HadGEM1, and (h) MRI.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Over tropical South America (15°S–5°N, 70°–45°W), the parent CMIP3 simulations indicate a range of possible changes in precipitation, from wetter conditions (CCSM3) to drier conditions (HadCM3). While the exact spatial distribution of the patterns differ, the sign of the precipitation change (future compared to twentieth century) between the parent CMIP3 simulations and our CAM–CLM-CN simulations is fairly consistent and seems to scale between the CMIP3 and CAM–CLM-CN simulations. This is demonstrated in Fig. 9, which shows a simple scatterplot of the CAM–CLM-CN precipitation anomalies (annual average future minus present) averaged over tropical South America compared to their CMIP3 parent anomalies (blue circles). With one exception, positive (negative) anomalies in the parent CMIP3 simulation are translated to positive (negative) anomalies in our CAM–CLM-CN simulations. However, there is a shift such that the CAM–CLM-CN simulations are wetter overall in the future than are their CMIP3 parents. To confirm this relationship, we also examined another tropical region, western and central Africa (0°–12°N, 15°W–15°E). The parent CMIP3 simulations indicate a range of possible changes in precipitation, from wetter conditions (CCSM3) to drier conditions (GFDL). All of our CAM–CLM-CN simulations indicate wetter conditions in the future (cf. Fig. 7 to Fig. 8). However, the degree to which our CAM–CLM-CN simulations become wetter in the future is modulated by the applied CMIP3 SSTs. This is demonstrated by the red circles in Fig. 9, which show that a wetter future in the CMIP3 simulations translates to a much wetter future in the CAM–CLM-CN simulations. A drier future in the CMIP3 simulations translates to a wetter future in the CAM–CLM-CN simulations, but to a lesser extent.

Scatterplot of CAM–CLM-CN precipitation anomalies (annual average future minus present; mm day−1) compared to their CMIP3 parent anomalies. Tropical South America is shown in blue (15°S–5°N, 70°–45°W) and western and central Africa (0°–12°N, 15°W–15°E) is shown in red.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Scatterplot of CAM–CLM-CN precipitation anomalies (annual average future minus present; mm day−1) compared to their CMIP3 parent anomalies. Tropical South America is shown in blue (15°S–5°N, 70°–45°W) and western and central Africa (0°–12°N, 15°W–15°E) is shown in red.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Scatterplot of CAM–CLM-CN precipitation anomalies (annual average future minus present; mm day−1) compared to their CMIP3 parent anomalies. Tropical South America is shown in blue (15°S–5°N, 70°–45°W) and western and central Africa (0°–12°N, 15°W–15°E) is shown in red.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
When considering the tropics as a whole (20°S–20°N), the ensemble average of our CAM–CLM-CN simulations shows increasing precipitation through the twenty-first century in the annual average (Fig. 10a, solid black line), whereas the parent CMIP3 ensemble indicates little trend (Fig. 10a, solid blue line). However, there are important seasonal variations to these tendencies. Our simulations do capture the drying during the Southern Hemisphere dry season (May–September) (Fig. 10b; cf. solid black line to solid blue line), which is part of an intensification of the annual cycle (wetter wet season and drier dry season) in future climate projections that is stronger in the Southern Hemisphere (Tan et al. 2008; Seth et al. 2011, 2013). Overall, these results indicate that CAM–CLM-CN does respond to the imposed SST patterns, but it is heavily modulated by the tendency of CAM–CLM-CN to favor a wetter future in tropical regions. Nonetheless, the wintertime drying that is critical for fire occurrence and vegetation change in the real world (e.g., Marengo et al. 2008; Lewis et al. 2011) is present, and this drying over tropical South America does prompt some interesting vegetation responses. These are outlined in section 4b(1). We will return to possible reasons for CAM–CLM-CN’s wet future projections in section 4b(2).

Mean and range of precipitation change (mm day−1) for CAM–CLM-CN runs (black/gray) and CMIP3 runs (blue) for (a) annual average over tropical land areas (20°S–20°N) and (b) Southern Hemisphere (5°–20°S) dry season (MJJAS). All differences are relative to 1970–99 means, computed for all models separately. Comparison of the solid blue and black lines shows the overall trend of the ensemble mean of the CMIP3 simulations vs the CAM–CLM-CN simulations, respectively.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Mean and range of precipitation change (mm day−1) for CAM–CLM-CN runs (black/gray) and CMIP3 runs (blue) for (a) annual average over tropical land areas (20°S–20°N) and (b) Southern Hemisphere (5°–20°S) dry season (MJJAS). All differences are relative to 1970–99 means, computed for all models separately. Comparison of the solid blue and black lines shows the overall trend of the ensemble mean of the CMIP3 simulations vs the CAM–CLM-CN simulations, respectively.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Mean and range of precipitation change (mm day−1) for CAM–CLM-CN runs (black/gray) and CMIP3 runs (blue) for (a) annual average over tropical land areas (20°S–20°N) and (b) Southern Hemisphere (5°–20°S) dry season (MJJAS). All differences are relative to 1970–99 means, computed for all models separately. Comparison of the solid blue and black lines shows the overall trend of the ensemble mean of the CMIP3 simulations vs the CAM–CLM-CN simulations, respectively.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
b. Projected vegetation change
1) Tropics
Figure 11 shows the ensemble mean changes in PFTs for 2070–99 compared to 1970–99 in the CAM–CLM-CNDV experiments. In tropical regions, there are modest increases in the coverage of the broadleaf evergreen PFT. These mainly occur on the poleward edges of their twentieth-century modeled distribution (e.g., over southern Brazil and central Africa). Increases in deciduous trees also occur over western Africa. These increases in tropical broadleaf and deciduous trees correspond to decreases in grasses and temperate needleleaf evergreens. The latter occurs spuriously in the control run, and the future change is due to exceedance of the maximum temperature limit for the temperate needleleaf evergreen of 18.8°C (Levis et al. 2004). Despite the use of different SST patterns, the sign and spatial extent of the vegetation changes are consistent among the simulations (not shown). However, a few simulations do show small decreases in broadleaf evergreen and deciduous tree cover over the tropics (20°S–20°N). We will explore these decreases in section 4b(2).

Ensemble average changes in PFTs (%) comparing the late twenty-first century (2070–99) and the twentieth century (1970–99) in CAM–CLM-CNDV simulations for (a) broadleaf evergreen trees, (b) needleleaf evergreen trees, (c) deciduous trees, (d) shrubs, (e) grasses, and (f) bare ground.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Ensemble average changes in PFTs (%) comparing the late twenty-first century (2070–99) and the twentieth century (1970–99) in CAM–CLM-CNDV simulations for (a) broadleaf evergreen trees, (b) needleleaf evergreen trees, (c) deciduous trees, (d) shrubs, (e) grasses, and (f) bare ground.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Ensemble average changes in PFTs (%) comparing the late twenty-first century (2070–99) and the twentieth century (1970–99) in CAM–CLM-CNDV simulations for (a) broadleaf evergreen trees, (b) needleleaf evergreen trees, (c) deciduous trees, (d) shrubs, (e) grasses, and (f) bare ground.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
As noted above, changes in mean vegetation cover over the tropics are fairly small; the fractional coverage of broadleaf evergreen trees and deciduous trees increases slightly from about 23% to 25.5% and from 21.5% to 23% (a percentage change of 11% and 7% of the original coverage, respectively) throughout the twenty-first century in the ensemble average. A few simulations show larger increases in broadleaf evergreen trees; these are the simulations forced by ECHAM5 and GISS. Interestingly, both of these models warm less in the eastern Pacific compared to their zonal mean warming (Fig. 2). Another model, CCSM3, also shows this departure but it does have more pronounced warming on the equator in the central Pacific. Lyon and Barnston (2005) showed that a robust relationship exists between the spatial extent of drought in the tropics and El Niño strength (based on Niño-3.4 SST anomalies); patterns that lack this characteristic are probably more likely to be wetter overall in the tropics. Indeed, the GISS-driven simulation has the largest trend in precipitation over land areas in the tropics [0.52 mm day−1 (95 yr)−1], while ECHAM is tied for the third-largest increase [0.40 mm day−1 (95 yr)−1]. In contrast to these two simulations, in the HadCM3-driven simulation there are a few sharp declines in broadleaf evergreen cover and deciduous tree cover (Fig. 12, red line). Because each event is followed by a fast recovery, the changes in vegetation cover are not due to the slow 20-yr climate envelope that can change PFT coverage in CAM–CLM-CNDV. Instead, these temporary decreases in tree cover result from the activation of the fire algorithm in CLM (described above in section 2).

Time series of vegetation cover (%) in the tropics from 2006–2100 in CAM–CLM-CNDV simulations for (a) broadleaf evergreen trees and (b) deciduous trees. The twenty-first-century ensemble mean is shown in dark gray and individual ensemble members are shown in light gray. The HadCM3-driven CAM–CLM-CNDV ensemble member is in red.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Time series of vegetation cover (%) in the tropics from 2006–2100 in CAM–CLM-CNDV simulations for (a) broadleaf evergreen trees and (b) deciduous trees. The twenty-first-century ensemble mean is shown in dark gray and individual ensemble members are shown in light gray. The HadCM3-driven CAM–CLM-CNDV ensemble member is in red.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Time series of vegetation cover (%) in the tropics from 2006–2100 in CAM–CLM-CNDV simulations for (a) broadleaf evergreen trees and (b) deciduous trees. The twenty-first-century ensemble mean is shown in dark gray and individual ensemble members are shown in light gray. The HadCM3-driven CAM–CLM-CNDV ensemble member is in red.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Figure 13a shows the fractional area burned over the tropics (20°S–20°N). From 2005 to the end of the century, fractional area burned increases by about 30% in the ensemble mean, and several large events are noticeable in a few ensemble members, including the HadCM3-driven ensemble member, which is shown in red. These large events are also visible in a frequency histogram of fractional area burned (Fig. 13b), comparing the eight simulations in the last 40 years of the twenty-first century to the first 40 years of the twenty-first century. Note that we compare the late twenty-first century to the early twenty-first century so that we can use all eight ensemble members so as to maintain the same number of values for each period. There is clearly an increase in the percentage of time a grid cell experiences a large fractional area burned (Fig. 13b). This relationship is maintained even when the HadCM3 ensemble member is removed (Fig. 13b, dashed lines). Over the tropics as a whole, fire season increases by about one week in the ensemble mean (7.21 days) over the twenty-first century (Fig. 14a).

(a) Fractional area burned (fraction) over the tropics (20°S–20°N) for CAM–CLM-CNDV twentieth-century control run (black), twenty-firs- century ensemble mean (dark gray), and individual ensemble members (light gray). The HadCM3-driven CAM–CLM-CNDV ensemble member is in red. (b) Frequency histogram of fractional area burned (fraction) for 2010–49 (blue) vs 2060–99 (yellow) considering all land grid points from 20°S–20°N for all ensemble members (solid lines) and excluding the HadCM3 forced ensemble member (dashed lines).
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

(a) Fractional area burned (fraction) over the tropics (20°S–20°N) for CAM–CLM-CNDV twentieth-century control run (black), twenty-firs- century ensemble mean (dark gray), and individual ensemble members (light gray). The HadCM3-driven CAM–CLM-CNDV ensemble member is in red. (b) Frequency histogram of fractional area burned (fraction) for 2010–49 (blue) vs 2060–99 (yellow) considering all land grid points from 20°S–20°N for all ensemble members (solid lines) and excluding the HadCM3 forced ensemble member (dashed lines).
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
(a) Fractional area burned (fraction) over the tropics (20°S–20°N) for CAM–CLM-CNDV twentieth-century control run (black), twenty-firs- century ensemble mean (dark gray), and individual ensemble members (light gray). The HadCM3-driven CAM–CLM-CNDV ensemble member is in red. (b) Frequency histogram of fractional area burned (fraction) for 2010–49 (blue) vs 2060–99 (yellow) considering all land grid points from 20°S–20°N for all ensemble members (solid lines) and excluding the HadCM3 forced ensemble member (dashed lines).
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Length of fire season (days) in the twentieth-century control run (black), twenty-first-century ensemble mean (dark gray), and individual ensemble members (light gray) in CAM–CLM-CNDV simulations over (a) the tropics (20°S–20°N) and (b) tropical South America (15°S–5°N, 70°–45°W). In both panels, the HadCM3-driven CAM–CLM-CNDV ensemble member is shown in red.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Length of fire season (days) in the twentieth-century control run (black), twenty-first-century ensemble mean (dark gray), and individual ensemble members (light gray) in CAM–CLM-CNDV simulations over (a) the tropics (20°S–20°N) and (b) tropical South America (15°S–5°N, 70°–45°W). In both panels, the HadCM3-driven CAM–CLM-CNDV ensemble member is shown in red.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Length of fire season (days) in the twentieth-century control run (black), twenty-first-century ensemble mean (dark gray), and individual ensemble members (light gray) in CAM–CLM-CNDV simulations over (a) the tropics (20°S–20°N) and (b) tropical South America (15°S–5°N, 70°–45°W). In both panels, the HadCM3-driven CAM–CLM-CNDV ensemble member is shown in red.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
These changes in dry season length are linked to an enhanced dry season in the future simulations, as indicated by the precipitation Hovmöller diagram in Fig. 15a. Figure 15a shows the precipitation climatology from the control simulation averaged over land areas over all longitudes in black contours. The annual cycle of precipitation clearly follows the solar annual cycle, reaching its maximum northward (southward) extension in the Northern (Southern) Hemisphere summer. The differences between the twenty-first and twentieth centuries (shaded) show an amplification of the annual cycle (i.e., wetter rainy seasons and drier dry seasons), a robust feature of precipitation change via the remote effect in future simulations (Tan et al. 2008; Seth et al. 2011, 2013). These decreases in winter precipitation are also apparent in Fig. 10b. These decreases in precipitation are reflected by changes in soil moisture, a variable which effectively integrates changes in precipitation, temperature, and evapotranspiration (Koster et al. 2004; Steiner et al. 2009). Recall that fire season length is controlled in part by near-surface soil moisture conditions in the fire module in CLM (Oleson et al. 2010). As a result of decreases in precipitation and increases in temperature (not shown), soil moisture decreases in the winter (dry) season of each hemisphere (Fig. 15b), leading to a lengthening of the fire season as shown in Fig. 14a.

(a) Hovmöller diagram of precipitation for the control simulation (black line; mm day−1) and differences between the late twenty-first century (2070–99) and the twentieth century (1970–99) (shaded) in the CAM–CLM-CNDV simulations over land areas. (b) As in (a), but for the top 0.5 m soil moisture (m3 m−3).
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

(a) Hovmöller diagram of precipitation for the control simulation (black line; mm day−1) and differences between the late twenty-first century (2070–99) and the twentieth century (1970–99) (shaded) in the CAM–CLM-CNDV simulations over land areas. (b) As in (a), but for the top 0.5 m soil moisture (m3 m−3).
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
(a) Hovmöller diagram of precipitation for the control simulation (black line; mm day−1) and differences between the late twenty-first century (2070–99) and the twentieth century (1970–99) (shaded) in the CAM–CLM-CNDV simulations over land areas. (b) As in (a), but for the top 0.5 m soil moisture (m3 m−3).
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
2) Tropical South America
Over tropical South America, fires with sufficient fractional area burned to remove vegetation (Fig. 16a, where the HadCM3-driven simulation is shown in red) occur in some of the simulations. Why do these changes happen in the simulations? First, large droughts that exceed observed z scores emerge in the twenty-first century in several of the simulations. Figure 17a shows the area-averaged precipitation over tropical South America for the twentieth and twenty-first centuries. All are calculated relative to the twentieth-century simulation (except observations). Noticeable observed droughts occur over tropical South America, with precipitation anomalies that exceed −1 standard deviation. Severe droughts occur in the HadCM3-forced simulation (red line) that exceed −2 standard deviations from the mean.

Fractional area burned (fraction) over tropical South America (5°S–5°N, 70°–45°W) for CAM–CLM-CNDV twentieth-century control run (black), twenty-first-century ensemble mean (dark gray), individual twenty-first century ensemble members (light gray), and HadCM3-driven ensemble member (red).
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Fractional area burned (fraction) over tropical South America (5°S–5°N, 70°–45°W) for CAM–CLM-CNDV twentieth-century control run (black), twenty-first-century ensemble mean (dark gray), individual twenty-first century ensemble members (light gray), and HadCM3-driven ensemble member (red).
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Fractional area burned (fraction) over tropical South America (5°S–5°N, 70°–45°W) for CAM–CLM-CNDV twentieth-century control run (black), twenty-first-century ensemble mean (dark gray), individual twenty-first century ensemble members (light gray), and HadCM3-driven ensemble member (red).
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Time series of surface climatic variables over tropical South America (15°S–5°N, 70°–45°W). (a) Precipitation z scores (distance from mean in standard deviation units) for CAMS–OPI observations (black solid line), twentieth-century CAM–CLM-CNDV control run (black dashed line), twenty-first-century CAM–CLM-CNDV ensemble mean (dark gray), twenty-first-century individual ensemble members (light gray), and CAM–CLM-CNDV HadCM3 ensemble member (red). (b) Top 0.5-m soil moisture (m3 m−3), (c) evapotranspiration (mm day−1), (d) surface net radiation (W m−2). (b)–(d) coloring as in (a), except no observations are shown.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Time series of surface climatic variables over tropical South America (15°S–5°N, 70°–45°W). (a) Precipitation z scores (distance from mean in standard deviation units) for CAMS–OPI observations (black solid line), twentieth-century CAM–CLM-CNDV control run (black dashed line), twenty-first-century CAM–CLM-CNDV ensemble mean (dark gray), twenty-first-century individual ensemble members (light gray), and CAM–CLM-CNDV HadCM3 ensemble member (red). (b) Top 0.5-m soil moisture (m3 m−3), (c) evapotranspiration (mm day−1), (d) surface net radiation (W m−2). (b)–(d) coloring as in (a), except no observations are shown.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Time series of surface climatic variables over tropical South America (15°S–5°N, 70°–45°W). (a) Precipitation z scores (distance from mean in standard deviation units) for CAMS–OPI observations (black solid line), twentieth-century CAM–CLM-CNDV control run (black dashed line), twenty-first-century CAM–CLM-CNDV ensemble mean (dark gray), twenty-first-century individual ensemble members (light gray), and CAM–CLM-CNDV HadCM3 ensemble member (red). (b) Top 0.5-m soil moisture (m3 m−3), (c) evapotranspiration (mm day−1), (d) surface net radiation (W m−2). (b)–(d) coloring as in (a), except no observations are shown.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
These HadCM3 droughts are associated with positive SST anomalies over the tropical North Atlantic, less warm anomalies over the tropical South Atlantic, and El Niño–like conditions in the tropical Pacific, as shown by the instantaneous correlation between area-averaged precipitation over tropical South America and observed SSTs (Figs. 18a,b) for the CAMS–OPI observations and CAM–CLM-CN control run, respectively. Note that these droughts occur in the HadCM3-driven CAM–CLM-CN and CAM–CLM-CNDV simulations in the same years (2065, 2097, and 2098; Figs. 18c–e), confirming that the SST anomalies are driving these events. Decreases in soil moisture contribute to the severity of these droughts. Despite an overall positive trend in annual average precipitation in the ensemble mean, soil moisture does not increase over the twentieth century over tropical South America (Fig. 17b). Consequently, fire season length increases in length by 2 weeks in the ensemble mean over the 95-yr simulation period (Fig. 14b). These changes are particularly notable in the HadCM3-driven CAM–CLM-CNDV ensemble member, shown in red, which shows the large droughts (Fig. 17a), large decreases in soil moisture (Fig. 17b), and marked decreases in evapotranspiration (Fig. 17c).

Correlation between annual average observed SSTs (°C) and precipitation (mm day−1) over tropical South America (15°S–5°N, 70°–45°W) showing (a) the correlation with CAMS–OPI observations and (b) the correlation with the CAM–CLM-CN control run precipitation from 1980 to 2004. Annual average SST anomalies (°C) for (c) 2065, (d) 2097, and (e) 2098 from the HadCM3 coupled simulation compared to the 1970–99 HadCM3 average.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1

Correlation between annual average observed SSTs (°C) and precipitation (mm day−1) over tropical South America (15°S–5°N, 70°–45°W) showing (a) the correlation with CAMS–OPI observations and (b) the correlation with the CAM–CLM-CN control run precipitation from 1980 to 2004. Annual average SST anomalies (°C) for (c) 2065, (d) 2097, and (e) 2098 from the HadCM3 coupled simulation compared to the 1970–99 HadCM3 average.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Correlation between annual average observed SSTs (°C) and precipitation (mm day−1) over tropical South America (15°S–5°N, 70°–45°W) showing (a) the correlation with CAMS–OPI observations and (b) the correlation with the CAM–CLM-CN control run precipitation from 1980 to 2004. Annual average SST anomalies (°C) for (c) 2065, (d) 2097, and (e) 2098 from the HadCM3 coupled simulation compared to the 1970–99 HadCM3 average.
Citation: Journal of Climate 28, 20; 10.1175/JCLI-D-14-00528.1
Returning to why annual average precipitation increases in the future in our simulations—and why our simulations produce wetter futures than their CMIP3 parents—recall the local and remote effects discussed in the introduction. In the remote response, the rising heat content of the ocean and higher SSTs result in nearly uniform warming in the upper troposphere (Sobel et al. 2001), increasing stability. Precipitation is reduced over tropical land areas where there is insufficient moist static energy to overcome this enhanced stability (Chou and Neelin 2004; Chou et al. 2009; Giannini 2010). In the local response, increases in net surface radiation lead to increases in evaporation and low-level moist static energy, driving increased precipitation and moisture convergence. In our simulations, the local effect appears to dominate since precipitation, net surface radiation, and evapotranspiration all increase on an annual average basis over tropical South America in nearly all ensemble members (Figs. 17a,c,d). Whether the remote or local effect is more important is likely linked to the strength of land surface–atmosphere interactions and therefore to the formulation of the land surface model, boundary layer, and convection parameterizations and deserves further exploration.
5. Discussion and conclusions
We have performed a series of experiments with CAM–CLM-CN and CAM–CLM-CNDV with SST warming patterns provided by eight of the CMIP3 projections under a medium–high emissions scenario. These experiments were designed to simulate a range of future climate–vegetation trajectories over the tropics. Because of the tendency of CAM–CLM-CN to favor the local response to climate change—that is, changes in net radiation at the surface are translated to increased evaporation, net surface radiation, and precipitation—overall we did not see as large a spread in our projected future precipitation on an annual average basis as in the original CMIP3 archive that provided the SST boundary conditions. In other words, the different SST warming patterns do have an impact, but this effect is modulated by CAM–CLM-CN. However, our simulations did faithfully reproduce the drier dry season present in the CMIP3 simulations, which has important implications for vegetation change.
In our simulations with dynamic vegetation enabled, the fractional coverage of broadleaf evergreen trees and deciduous trees over the tropics increases from about 23% to 25.5% (a percentage difference of 11%) and from 21.5% to 23% (7% increase), respectively, throughout the twenty-first century in the ensemble average. An exception is the HadCM3-driven experiment, which experiences several sharp declines and recoveries in tree cover over the tropics. As a result of the enhanced dry season, longer fire seasons and increases in fractional area burned are found over the tropics as a whole and in particular over tropical South America. These changes are evident even when the HadCM3-forced ensemble member is excluded. Our results echo many other studies of vegetation change over the region. Enhanced fire risk was also found in a study using the vegetation–global–atmosphere–soil (VEGAS) model driven by output from 24 CMIP3 models (Cook et al. 2012). Similar to our results, Cook et al. (2012) found that a drier dry season (May–September) projected by the future scenario simulations leads to lower soil moisture and heightened fire risk in the Amazon. A recent study by Galbraith et al. (2010) suggests that DGVMs may in fact be underestimating the effects of soil moisture stress on vegetation based on field experiments where throughfall reaching the soil was reduced, leading to reduced biomass.
In the HadCM3-forced ensemble member, extreme drought occurrences over tropical South America associated with El Niño–like warming patterns in the Pacific and a meridional gradient in tropical Atlantic SSTs lead to fires that remove all vegetation cover over the eastern Amazon. While the other ensemble members show droughts and fire pulses as well, the HadCM3-forced result appears to be unique in its extreme response. Modeling studies that use HadCM3 climate forcing typically find decreases in biomass due to the HadCM3 projected decrease in precipitation. For example, Sitch et al. (2008) found that five different dynamic vegetation models [hybrid land terrestrial ecosystem (HyLand), ORCHIDEE, Sheffield-DVGM (SHE), TRIFFID, and LPJ] simulate decreases in vegetation carbon over Amazonia when the models are forced with the HadCM3 data. Huntingford et al. (2013) investigated how biomass stocks would change using the MOSES–TRIFFID land surface model forced with climate data from 22 of the CMIP3 models. Only in the HadCM3-forced simulation did biomass decrease over the tropical Americas. Malhi et al. (2009) calculated accumulated water stress from CMIP3 output for the twentieth and twenty-first centuries and found that there is a high probability of a more intense dry season in the future. Water stress increases over eastern Amazonia to the point where the region crosses a threshold to transition to savanna when the output from HadCM3 is used. Overall, their study predicts a climate over the eastern Amazon more suitable to seasonal forest, with the potential for enhanced fire risk.
The relative importance of the climate forcing and the physiological response is still uncertain. The similarities in responses across multiple DGVMs to the HadCM3 forcing suggest that the climate forcing is the most important factor when determining future trajectories of vegetation. However, if the fertilization response to CO2 is strong, biomass could increase despite precipitation decreases (Rammig et al. 2010). Substantial structural uncertainties remain in both our understanding of vegetation processes such as mortality events (McDowell et al. 2013) and how to model them (Adams et al. 2013). Our results and existing studies suggest that a large-scale reduction in forest cover is an unlikely but still possible event given the resemblance of recent droughts to CMIP3 HadCM3 projection (Marengo et al. 2008; Lewis et al. 2011). Similar to Malhi et al. (2009), we find that the risk is likely greatest in seasonally dry forests given the amplification of the precipitation annual cycle and drier springs simulated here in agreement with both CMIP3 (Seth et al. 2011) and CMIP5 (Seth et al. 2013) projections. This is in contrast to our results for the midlatitudes, where heat stress was the primary factor behind vegetation change (Jiang et al. 2013) in partial agreement with observed relationships (Williams et al. 2013).
Acknowledgments
We thank three anonymous reviewers for their comments, which greatly helped to improve the content and presentation of this manuscript. We thank A. T. Hoang and S. Levis for their help with obtaining CMIP3 data and CLM4 initial data files. We thank S. Levis, D. Lawrence, and A. Giannini for helpful discussions. Los Alamos National Laboratory LDRD and DOE Office of Science (BER) provided funding for this project. The National Center for Atmospheric Research (NCAR) is operated by the University Corporation for Atmospheric Research under sponsorship of the National Science Foundation.
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See also Fig. 1 in Seth et al. (2013).